460ACANTHACEAEovate, (2–)3–5.5 × 1.5–2.5 cm, variable in size, papery, abaxiallylight green and pilose along midvein, adaxially green andpilose especially along veins, base broadly cuneate and decurrentonto petiole, margin entire, apex shortly acuminate to acute.Spikes terminal, 5–7(–10) cm, unbranched, nodes distant, with2–4 flowers per node; peduncle short; rachis densely pilose;bracts linear-lanceolate, 3–5 cm; bracteoles similar to bracts.Calyx ca. 7 mm, 5-lobed almost to base; lobes linear-lanceolate,5–6 mm, pilose along midvein. Corolla white, 1–1.3 cm, outsidesomewhat pubescent to glabrescent; tube short; limb longerthan tube; lower lip 4–5 mm, 3-lobed, lobes pilose inside andciliate on margin; upper lip 2-lobed. Staminal filaments pilose;anther thecae parallel, superposed, lower one with a white spurat base. Ovary glabrous; style densely pilose. Capsule ca. 2 cm,4-seeded. Seeds reddish brown, circular in outline, 3–4 mm indiam., rugose-verruculose. Fl. Jun–Aug.● Thickets; 800–1200 m. Sichuan, Yunnan.37. Justicia damingensis (H. S. Lo) H. S. Lo, Guihaia 17: 51.1997.大 明 爵 床 da ming jue chuangMananthes damingensis H. S. Lo, Bull. Bot. Res., Harbin1(4): 106. 1981.Herbs 15–30 cm tall. Stems subterete, basally prostrateand rooting at nodes then erect, sulcate, bifariously pubescent.Petiole 0.5–1.5 cm, pubescent; leaf blade ovate, 4–9 × 2.5–4.5cm, thinly papery, abaxially pubescent along veins, adaxiallyglabrous, secondary veins ca. 5 on each side of midvein, basenarrowly cuneate, margin shallowly undulate, apex acute. Racemesterminal, spiciform, 17–18 cm, branchless or trifurcatebranched, with 3 flowers per node; peduncle 5.5–10 cm, pubescent;bracts subulate, 1.5–2 mm, abaxially pubescent; bracteolessimilar to bracts. Pedicel ca. 1 mm, pubescent. Calyx ca. 6mm, 5-lobed almost to base; lobes narrowly triangular, ca. 5 ×0.6 mm, margin ciliate, apex acuminate. Corolla yellow, ca. 1cm, outside puberulent; lower lip ca. 3.2 mm, patent, 3-lobed,lobes ovate and apex rounded; upper lip subdeltoid, ca. 2.5 mm.Staminal filaments ca. 3 mm, glabrous; anther thecae ellipsoid,superposed, lower one spurred at base. Ovary glabrous; styleca. 6 mm, glabrous. Capsule not seen. Fl. May–Jun.● Thickets; 300–600 m. Guangxi.38. Justicia amblyosepala D. Fang & H. S. Lo, Guihaia 17:55. 1997.钝 萼 爵 床 dun e jue chuangMananthes amblyosepala (D. Fang & H. S. Lo) C. Y. Wu& C. C. Hu.Shrubs 30–50 cm tall. Stems terete, glabrous. Petiole 0.8–3cm, glabrous; leaf blade ovate to elliptic, 9–17 × 4.5–8 cm, papery,glabrous, secondary veins 5–7 on each side of midvein,base attenuate to rounded, margin subentire, apex acute. Spikesterminal or axillary, 2–23 cm, interrupted, simple or branched;peduncle 1.5–5 cm; rachis 4-angled, sulcate, bifariously puberulent,with 1 flower per node; bracts triangular to rarely narrowlytriangular, 1.5–3 mm, subglabrous; bracteoles narrowlytriangular, ca. 2 mm, margin ciliolate. Calyx ca. 5 mm, 5-lobed;lobes elliptic-lanceolate, both surfaces usually puberulent tosubglabrous, apex obtuse to rounded. Corolla light yellow, ca.1.3 cm, outside retrorsely puberulent; lower lip 3-lobed, lobesca. 1.5 mm and apex rounded; upper lip ovate-triangular, ca. 5mm, erect, apex emarginate. Staminal filaments ca. 3 mm, glabrous;anther thecae ellipsoid, ca. 1.5 mm, unequal, superposed,lower one spurred at base. Ovary pubescent at tip; style 8–11mm, sparsely puberulent. Capsule clavate, ca. 1.5 cm, pubescent.Seeds ovate in outline, ca. 1 × 1 mm, verrucose. Fl. Mar–May, fr. Aug.● Forests on limestone slopes. Guangxi.39. Justicia acutangula H. S. Lo & D. Fang, Guihaia 17: 56.1997.棱 茎 爵 床 leng jing jue chuangMananthes acutangula (H. S. Lo & D. Fang) C. Y. Wu &C. C. Hu.Herbs 50–100 cm tall. Stems sharply 4-angled and slightlywinged, sulcate, glabrous. Petiole ca. 1.3 mm, pubescent; leafblade elliptic to rarely broadly elliptic, 8.5–28 × 5–12.5 cm,glabrous, secondary veins 13–15 on each side of midvein, baserounded to attenuate and decurrent onto petiole, margin subentire,apex acute. Spikes terminal or axillary in upper leaf axils,3.5–14 cm, interrupted, unbranched or branched, with 1 flowerper node; bracts triangular, 2–3 mm, glabrous, margin ciliate;bracteoles ca. 2.5 mm, glabrous, margin ciliate. Calyx ca. 5mm, outside densely puberulent, 5-lobed almost to base; lobeslanceolate, apex acute. Corolla light greenish yellow with purplishdots on lower lip, 2–2.2 cm, outside pubescent; limb aslong as tube; lower lip as long as upper lip, 3-lobed, lobesovate, ca. 2 mm, and apex rounded; upper lip ca. 9 mm, apexemarginate. Staminal filaments ca. 7.5 mm, glabrous; antherthecae ca. 2.5 mm, superposed, lower one minutely spurred atbase. Ovary glabrous; style ca. 2 cm, basal portion pubescent.Capsule clavate, ca. 2 cm. Seeds ovate in outline, ca. 1.2 × 1.3mm. Fl. Nov–Jan, fr. Apr–May.● Forests on limestone hills; 500–700 m. Guangxi, Guizhou.40. Justicia patentiflora Hemsley, Hooker’s Icon. Pl. 28: t.2792. 1905.野 靛 棵 ye dian keMananthes patentiflora (Hemsley) Bremekamp.Herbs to 2 m tall, perennial. Stems subterete, glabrous.Petiole 2–6 cm; leaf blade ovate to oblong-lanceolate, 16–26 ×7.5–9.5 cm, membranous, glabrous, secondary veins 8 or 9 eachside, blade basally attenuate to a narrowly cuneate base, apexacuminate. Spikes terminal or axillary in upper leaf axils, 5–12cm, unbranched or bifurcately branched; rachis 4-angled, bifariouslyyellowish pubescent, with 1 flower per node; bracts scalelike,triangular-lanceolate, ca. 3 mm, abaxially puberulent;bracteoles similar to bracts. Calyx ca. 1.2 cm, puberulent, 5-lobed almost to base; lobes linear, apex acute. Corolla purplishred with purplish spots, 1.5–2 cm; tube 1–1.5 cm, abruptlyrecurved above base, outside puberulent; lower lip 3-lobed,lobes circular; upper lip subcircular, shorter than lower lip, apexemarginate. Staminal filaments ca. 5 mm, glabrous; anther
ACANTHACEAE 461thecae superposed, lower one spurred at base. Ovary glabrous;style ca. 7 mm, basal portion sparsely puberulent. Capsulenarrowly obovoid, ca. 1.5 cm. Seeds light yellow, circular inoutline, ca. 1.2 × 1.2 mm, verrucose. Fl. Nov–Jan, fr. Apr–May.Forests, streamsides, limestone hills; 500–1800(–2400) m. Yunnan[Vietnam].“Mananthes tubiflora” (Y. M. Shui & W. H. Chen, Seed Pl. KarstReg. China 1: 182. 2006) belongs here but was not validly published.41. Justicia poilanei Benoist, Notul. Syst. (Paris) 5: 125. 1936.毛 萼 爵 床 mao e jue chuangShrubs to 2 m tall. Stems ± 4-angled, sulcate, glabrous.Petiole 1–6 cm, flat, sulcate, margin ciliate; leaf blade ovate toovate-lanceolate, 10–20 × 4–8 cm, glabrous, secondary veins6–10 on each side of midvein, base attenuate and decurrent ontopetiole, margin entire or undulate, apex acute. Spikes terminalor rarely axillary, 7–15 cm, usually trifurcately branched orrarely unbranched; peduncle 1–3 cm; rachis 4-angled, glabrous,with 1 flower per node; bracts triangular, 2–2.5 × ca. 2 mm,both surfaces glabrous, margin ciliate; bracteoles similar tobracts. Calyx 3–4 mm, both surfaces puberulent especially distallyon inner surface, 5-lobed almost to base; lobes lanceolate,margin ciliate, apex acute. Corolla ca. 1 cm, outside pubescent;tube as long as limb; lower lip 3-lobed, lobes subequal, ovate,and apex rounded; upper lip subtriangular, erect, apex emarginate.Staminal filaments ca. 4 mm, glabrous; anther thecaeunequal, superposed, lower one spurred at base. Ovary glabrous;style ca. 8 mm, glabrous. Capsule not seen. Fl. Dec–Feb.Limestone hills; 1000–2400 m. Yunnan [Vietnam].42. Justicia kampotiana Benoist, Notul. Syst. (Paris) 5: 118.1936.那 坡 爵 床 na po jue chuangMananthes kampotiana (Benoist) C. Y. Wu & C. C. Hu.Shrubs to 1 m tall. Stems ± 4-angled, sulcate, glabrous.Petiole 3–6 mm, glabrous; leaf blade lanceolate, 9–16 × 4.3–5cm, glabrous, secondary veins ca. 8 on each side of midvein,base cuneate, margin entire, apex shortly acuminate. Spikesaxillary or terminal; rachis 4-angled, unbranched or 2-furcatelybranched, subglabrous, 1-flowered per node; bracts narrowlytriangular, ca. 1.5 mm, abaxially pubescent; bracteolessimilar to bracts. Calyx ca. 3 mm, outside pubescent, inside glabrous,5-lobed almost to base; lobes linear, subequal, apexacute. Corolla ca. 1 cm, outside pubescent; tube slightly longerthan limb; lower lip 3-lobed, lobes subequal, ovate, and apexrounded; upper lip subtriangular, erect, apex emarginate. Staminalfilaments ca. 4 mm, glabrous; anther thecae unequal, superposed,lower one spurred at base. Ovary and style glabrous.Capsule glabrous. Fl. Apr–May.On rocks in forests; 500–600 m. Guangxi [Cambodia].43. Justicia caudatifolia (H. S. Lo & D. Fang) Z. P. Hao, Y. F.Deng & T. F. Daniel, J. Trop. Subtrop. Bot. 18: 486. 2010.尾 叶 爵 床 wei ye jue chuangLeptostachya caudatifolia H. S. Lo & D. Fang, Guihaia17: 45. 1997.Herbs to 1 m tall. Stems 4-angled, basally prostrate androoting at nodes, sulcate, glabrous. Petiole 2–15 mm, glabrous;leaf blade elliptic to obovate-elliptic, 3–10 × 1.2–4.2 cm, apicalhalf broadest, papery, glabrous, secondary veins 5 or 6 on eachside of midvein, base cuneate, margin subentire, apex caudate.Spikes axillary, 1–3 cm, interrupted; peduncle 0.5–1.5 cm; rachissulcate, puberulent, 1-flowered per node; bracts linear-triangular,3–4 mm, abaxially puberulent, margin ciliate; bracteolesovate, 2.5–3 mm, margin ciliate. Calyx ca. 4 mm, 5-lobedalmost to base; lobes narrowly triangular, outside glabrescent,inside glabrous. Corolla white, ca. 1.1 cm, outside slightly retrorselypubescent; tube as long as limb; lower lip 3-lobed,lobes ca. 2.5 mm and apex rounded; upper lip ca. 5 mm, apexemarginate. Staminal filaments ca. 3 mm, glabrous; antherthecae superposed, lower one minutely spurred at base. Ovaryglabrous; style ca. 5 mm, glabrous. Capsule not seen. Fl. Oct.● On rocks in forests on limestone hills. Guangxi.25. RHINACANTHUS Nees in Wallich, Pl. Asiat. Rar. 3: 76, 108. 1832.灵 枝 草 属 ling zhi cao shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielShrubs, subshrubs, or herbs, with cystoliths. Leaves opposite, petiolate or subsessile; leaf blade margin entire or slightly undulate.Inflorescences axillary or terminal, spikes or racemes, sometimes branched into panicles; bracts and bracteoles shorter thancalyx. Calyx deeply 5-lobed; lobes subequal. Corolla white or greenish [or purple]; tube cylindric, elongate, narrow; limb 2-lipped;lower lip 3-lobed, lobes ascending cochlear in bud; upper lip entire or 2-cleft, inside rugulose. Stamens 2, exserted from mouth ofcorolla; anthers 2-thecous; thecae unequally inserted to superposed, lower theca with or without a basal appendage; staminodesabsent. Capsule with a solid stalk, clavate, 4-seeded; retinacula present. Seeds discoid, variously ornamented, lacking trichomes.About 25 species: tropical and subtropical regions of Africa and Asia; two species (one endemic) in China.Rhinacanthus calcaratus (Wallich) Nees was reported from Yunnan by C. C. Hu (FRPS 70: 268. 2002) based on a misidentification of Pseuderanthemumpolyanthum.1a. Leaf blade narrowly obovate to oblanceolate, 20–24 × 6–8 cm, abaxially glabrous, secondary veins 8–10 oneach side of midvein; corolla tube ca. 4 cm, lobes of lower lip 2–2.5 cm .................................................................. 1. R. beesianus1b. Leaf blade elliptic, ovate-elliptic, or rarely lanceolate, 2–7(–11) × 0.8–3 cm, abaxially pubescent, secondaryveins 5 or 6 on each side of midvein; corolla tube 1.5–1.8 cm, lobes of lower lip 2–4 mm ......................................... 2. R. nasutus
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ACANTHACEAE爵 床 科 jue chuang k
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ACANTHACEAE 37122b. Anthers 2-theco
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ACANTHACEAE 37311a. Inflorescences
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ACANTHACEAE 375lary or terminal, 2-
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ACANTHACEAE 377Two species: China,
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ACANTHACEAE 379Vines, herbaceous. S
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ACANTHACEAE 3817. STROBILANTHES Blu
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ACANTHACEAE 383Type 10. S. tetraspe
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ACANTHACEAE 38525b. Plants strongly
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ACANTHACEAE 38775a. Calyx lobes lin
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ACANTHACEAE 389115b. Corolla outsid
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ACANTHACEAE 391times rosette-formin
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ACANTHACEAE 393576. 1784; Acanthopa
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ACANTHACEAE 395with a whitish tube
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ACANTHACEAE 397Herbs 30-80 cm tall,
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ACANTHACEAE 399Strobilanthes petiol
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ACANTHACEAE 401silky tomentose and
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ACANTHACEAE 4036 × 1.2-1.5 mm. Cor
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ACANTHACEAE 405acute to shortly acu
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ACANTHACEAE 407Subshrubs to 20 cm t
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