448ACANTHACEAErolla throat; filaments 1.5–4 mm, glabrous or puberulent; antherthecae 1–1.5 mm. Ovary glabrous or apically hirsute; style glabrousor hirsute at basal portion; stigma 2-cleft. Capsule 1–1.5cm, finely pubescent. Seeds ca. 2 × 2 mm, with concentricridges of short linear warts. Fl. Jun–Sep.Moist forests; 900–1600 m. Guangdong, Guangxi, Hainan [Bhutan,India, Indonesia, Laos, Malaysia, Myanmar, Thailand, Vietnam].22. CODONACANTHUS Nees in A. Candolle, Prodr. 11: 103. 1847.钟 花 草 属 zhong hua cao shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielHerbs, small, erect, perennial, with cystoliths. Leaves petiolate; leaf blade margin entire to subsinuate. Inflorescences of axillaryor terminal racemes, loose, when terminal sometimes branched and forming a panicle, pedicellate; bracts and bracteoles minute.Flowers 1(–3 or more) in only 1 axil per node. Calyx deeply 5-lobed; lobes subequal. Corolla white with colored markings on lowerlip, broadly campanulate; tube short; limb slightly 2-liped; lower lip 3-lobed; upper lip 2-lobed; lobes ascending cochlear in bud.Stamens 2, included in to slightly emergent from corolla tube; anthers 2-thecous; thecae unequal, subparallel, ± equally to subequallyinserted; staminodes 2, short. Ovary with 2 ovules per locule; stigma capitate to shallowly 2-lobed. Capsule with a solid stalk at base,fertile region ellipsoid, usually with a medial constriction, up to 4-seeded; retinacula present. Seeds ± discoid, surfaces and marginsmooth to ± rugose, lacking trichomes.Two species: E and S Asia; one species in China.1. Codonacanthus pauciflorus (Nees) Nees in A. Candolle,Prodr. 11: 103. 1847.钟 花 草 zhong hua caoAsystasia pauciflora Nees in Wallich, Pl. Asiat. Rar. 3: 90.1832; Codonacanthus acuminatus Nees; Leptostachya repandaQ. H. Chen.Herbs 20–60 cm tall, much branched or unbranched. Stems4-angled, sulcate, 2-fariously pubescent. Petiole 3–15 mm,puberulent or glabrous; leaf blade elliptic, ovate, or lanceolate,(2–)4–12 × (0.5–)1.5–4.5 cm, thin, abaxially pale green andglabrous or sparsely pubescent on major veins, adaxially green,glabrous or with few large-celled hairs, and with numerouscystoliths, secondary veins 5–7 on each side of midvein, baserounded, cuneate, or attenuate and decurrent onto petiole, apexacute or acuminate. Racemes simple or terminal ones usuallyforming a panicle up to 15 cm; rachis puberulent; bracts lanceolateto linear, 1–2.5 mm, puberulent; bracteoles lanceolate,1–2 mm, puberulent. Pedicel 1–7 mm, puberulent. Calyx 2.5–4mm; lobes lanceolate, puberulent. Corolla white with purplespots at base of lower lip, 6.5–10 mm, glabrous; tube basallycylindric for 1–2 mm and 1–1.5 mm wide then abruptlywidened to 3.5–6.5 mm at mouth; lobes of lower lip 3.5–5.5 ×2–3.5 mm. Staminal filaments ca. 2.5 mm, glabrous; antherthecae 1.2–1.4 mm. Ovary glabrous; style 4–6.5 mm. Capsule1–1.9 cm, glabrous. Seeds 2–3 mm. Fl. and fr. Aug–Apr.Evergreen broad-leaved forests, wet places in ravines; below 100–1500 m. Fujian, Guangdong, Guangxi, Guizhou, Hainan, Jiangxi, Taiwan,Yunnan [Bhutan, Cambodia, India, Japan, Myanmar, Thailand,Vietnam].23. COSMIANTHEMUM Bremekamp, Blumea 10: 166. 1960.秋 英 爵 床 属 qiu ying jue chuang shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielHerbs, often prostrate then ascending, rarely erect, with cystoliths, isophyllous. Leaves petiolate or subsessile; leaf blade marginentire. Inflorescences terminal thyrsoid racemes or thyrses, sometimes with many-flowered cymes proximally and 1–3-floweredcymes distally; bracts small; bracteoles smaller than bracts. Calyx usually deeply 5-lobed; lobes subequal or slightly unequal, posteriorone sometimes shorter. Corolla white, greenish, or yellowish; tube straight or curved, sometimes with a dorsal pouch; throat shortor absent; limb 2-lipped; lower lip 3-lobed; upper lip emarginate or 2-lobed [or entire]; lobes ascending cochlear in bud. Stamens 2,inserted at throat; anthers 2-thecous; thecae equal or subequal, parallel, muticous at base; staminodes 2, inserted at base of upper lip.Stigma 2-cleft. Capsule stalked, 4-seeded; retinacula present.About ten species: SE Asia, especially Borneo; three species (two endemic) in China.Cosmianthemum resembles Pseuderanthemum in fruit and androecium but differs by the generally smaller corolla being distinctly 2-lipped (theupper lip only shallowly lobed) and lacking an elongate, narrow, cylindric tube; see B. Hansen (Nordic J. Bot. 5: 195–197. 1985) and B. L. Burtt andR. M. Smith (Notes Roy. Bot. Gard. Edinburgh 26: 365–381. 1965).1a. Leaf blade ovate to broadly ovate, 4–9 cm wide .................................................................................................. 1. C. viriduliflorum1b. Leaf blade elliptic-lanceolate, ovate-elliptic, or oblong-lanceolate, 1.4–7.3 cm wide.2a. Bracts abaxially gland-tipped pubescent; calyx outside gland-tipped pubescent ............................................ 2. C. knoxiifolium2b. Bracts abaxially non-gland-tipped pubescent; calyx outside non-glandular pubescent .................................. 3. C. guangxiense
ACANTHACEAE 4491. Cosmianthemum viriduliflorum (C. Y. Wu & H. S. Lo) H.S. Lo, Guihaia 17: 42. 1997.海 南 秋 英 爵 床 hai nan qiu ying jue chuangGraptophyllum viriduliflorum C. Y. Wu & H. S. Lo, Fl.Hainan. 3: 594. 1974.Herbs to 60 cm tall, erect, perennial. Stems terete, glabrous;branches yellowish brown, pubescent. Petiole 1–2.5 cm,pubescent; leaf blade ovate to broadly ovate, 7–15 × 4–9 cm,both surfaces glabrous, abaxially light green, adaxially darkgreen and lustrous, secondary veins 5–7 on each side of midveinand netted near margin, base broadly cuneate to subrounded,margin entire or inconspicuously undulate, apex acuminate,caudate, or sometimes acute. Thyrses terminal, 4–10cm; peduncle 1–4 cm; rachis pubescent; bracts linear, 1–1.5 ×ca. 0.5 mm; bracteoles linear, ca. 1 × 0.5 mm. Pedicel 2–2.5mm. Calyx ca. 5 mm, gland-tipped pubescent; lobes linear-lanceolate,apex acuminate. Corolla light greenish yellow, ca. 1.2cm, outside and inside pubescent; lobes of lower lip oblonglanceolate,ca. 3 × 0.6 mm, subacute at apex; upper lip erect,apex 2-cleft to emarginate. Staminal filaments ca. 4 mm, pubescent;anther thecae ca. 2 mm; staminodes ca. 2 mm, sparselypubescent. Ovary glabrous; style ca. 6 mm, sparsely pubescent.Capsule ca. 2 cm. Seeds not seen. Fl. Jul–Oct.● Forests; 700–1000 m. Hainan.2. Cosmianthemum knoxiifolium (C. B. Clarke) B. Hansen,Nordic J. Bot. 5: 195. 1985.秋 英 爵 床 qiu ying jue chuangGymnostachyum knoxiifolium C. B. Clarke, J. Asiat. Soc.Bengal, Pt. 2, Nat. Hist. 74: 663. 1908; Pseuderanthemumparviflorum Ridley; Ptyssiglottis parviflora (Ridley) Ridley;Sphinctacanthus malayanus Ridley.Shrubs to 50 cm tall or more. Stems terete, puberulent.Petiole 1–1.5 cm, glabrous; leaf blade elliptic-lanceolate, 8–20× 1.8–7.3 cm, leathery, abaxially puberulent along veins andmargin, adaxially glabrous, secondary veins 8–10 on each sideof midvein, base cuneate, margin remotely serrate or subentire,apex acuminate to attenuate. Thyrses terminal, (5.5–)10–30(–60) cm; rachis brownish pubescent with few gland-tippedtrichomes; bracts lanceolate, 2–3 × 0.5–1 mm, gland-tipped pubescent;bracteoles subulate, 1–2 × ca. 0.5 mm. Pedicels 1–3mm, pubescent. Calyx 2.5–5 mm, outside gland-tipped pubescent;lobes lanceolate, outside scaly hairy. Corolla white to paleyellow with purple spots on lower lip, ca. 1.5 cm, outside pubescent;tube ca. 5 mm; upper lip erect, apically 2-lobed toemarginate. Staminal filaments 6–7 mm, glabrous; staminodesca. 1 mm, pubescent. Ovary gland-tipped pubescent; style pubescent.Capsule 2–2.7 cm, puberulent. Seeds ovate in outline,ca. 2 × 2 mm, flattened, irregularly ridged or warty, lackingtrichomes. Fl. May–Sep, fr. Sep–Dec.Dense montane forests; 400–500 m. Hainan [Malaysia, Thailand,Vietnam].3. Cosmianthemum guangxiense H. S. Lo & D. Fang, Guihaia17: 42. 1997.广 西 秋 英 爵 床 guang xi qiu ying jue chuangCosmianthemum longiflorum D. Fang & H. S. Lo.Herbs to 70 cm tall, branched or unbranched. Stems 4-angled to subterete, sulcate, sparsely pubescent. Petiole 0.5–1cm, grooved, pilose; leaf blade ovate-elliptic to oblong-lanceolate,2–13.5 × 1.4–4 cm, abaxially glabrous, adaxially pilosetoward base or along veins, secondary veins 5–7 on each side ofmidvein, base rounded to cuneate, margin entire, apex acuminate.Thyrses terminal, 3–10 cm; rachis pilose; peduncle(0.5–)1–3 cm, pubescent; cymes 1–6-flowered; bracts triangular-subulate,ca. 1.5 × 0.5 mm, pilose; bracteoles similar tobracts. Pedicel 1–3 mm. Calyx 3–4 mm, outside pilose; lobeslinear, apex acute. Corolla white or yellowish green, 1–1.5 cm,outside sparsely pubescent to subglabrous, inside glabrous; tubeca. 1.5 mm wide for 4–5 mm; lower lip 6–7 × 3.5–5 mm, lobesovate and ca. 1 × 1 mm; upper lip ovate, 5–6 × ca. 5 mm.Staminal filaments 4–5 mm, glabrous; thecae ca. 2 mm. Ovaryglabrous; style 0.9–1.1 cm, sparsely hirsute to subglabrous.Capsule and seeds not seen. Fl. Oct.● Forests; ca. 400 m. Guangxi.24. JUSTICIA Linnaeus, Sp. Pl. 1: 15. 1753.爵 床 属 jue chuang shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielAdatoda Adanson; Adhatoda Miller; Calophanoides Ridley; Ecbolium Kuntze (1891), not Kurz (1871); Gendarussa Nees;Mananthes Bremekamp; Rhaphidospora Nees; Rostellaria Nees (1832), not C. F. Gaertner (1807); Rostellularia Reichenbach.Herbs, subshrubs, or shrubs, cystoliths present. Leaves sessile or petiolate; leaf blade margin usually entire but sometimessinuate or slightly serrate. Inflorescences dichasia (sometimes reduced to a single flower) in leaf axils or axillary or terminal spikes,racemes, or thyrses, sometimes branched and becoming panicles; bracts variable in shape, sometimes prominent and/or brightlycolored; bracteoles 2, similar to or smaller than bracts. Calyx deeply 4- or 5-lobed; lobes equal or subequal. Corolla tubular orfunnel-shaped; limb strongly 2-lipped; lower lip 3-lobed; upper lip entire to 2-lobed, internally rugulate (i.e., with a stylar furrow);lobes ascending cochlear in bud. Stamens 2; anthers 2-thecous; thecae equally or unequally inserted, parallel to perpendicular,one or both spurred at base or sometimes lacking basal appendages; staminodes absent. Ovary with 2 ovules per locule;stigma slightly 2-lobed. Capsule with a sterile basal stalk and a fertile head (i.e., ± clavate), 2–4-seeded; retinacula present. Seedscompressed [to spheric].About 700 species: tropical and temperate regions worldwide; 43 species (26 endemic, three introduced) in China.
- Page 1 and 2:
ACANTHACEAE爵 床 科 jue chuang k
- Page 3 and 4:
ACANTHACEAE 37122b. Anthers 2-theco
- Page 5 and 6:
ACANTHACEAE 37311a. Inflorescences
- Page 7 and 8:
ACANTHACEAE 375lary or terminal, 2-
- Page 9 and 10:
ACANTHACEAE 377Two species: China,
- Page 11 and 12:
ACANTHACEAE 379Vines, herbaceous. S
- Page 13 and 14:
ACANTHACEAE 3817. STROBILANTHES Blu
- Page 15 and 16:
ACANTHACEAE 383Type 10. S. tetraspe
- Page 17 and 18:
ACANTHACEAE 38525b. Plants strongly
- Page 19 and 20:
ACANTHACEAE 38775a. Calyx lobes lin
- Page 21 and 22:
ACANTHACEAE 389115b. Corolla outsid
- Page 23 and 24:
ACANTHACEAE 391times rosette-formin
- Page 25 and 26:
ACANTHACEAE 393576. 1784; Acanthopa
- Page 27 and 28:
ACANTHACEAE 395with a whitish tube
- Page 29 and 30: ACANTHACEAE 397Herbs 30-80 cm tall,
- Page 31 and 32: ACANTHACEAE 399Strobilanthes petiol
- Page 33 and 34: ACANTHACEAE 401silky tomentose and
- Page 35 and 36: ACANTHACEAE 4036 × 1.2-1.5 mm. Cor
- Page 37 and 38: ACANTHACEAE 405acute to shortly acu
- Page 39 and 40: ACANTHACEAE 407Subshrubs to 20 cm t
- Page 41 and 42: ACANTHACEAE 40910 × ca. 3 mm, apic
- Page 43 and 44: ACANTHACEAE 4111b. Stems, petioles,
- Page 45 and 46: ACANTHACEAE 413woody. Leaves subses
- Page 47 and 48: ACANTHACEAE 415gins white ciliolate
- Page 49 and 50: ACANTHACEAE 417long acuminate; brac
- Page 51 and 52: ACANTHACEAE 419retaining style; tub
- Page 53 and 54: ACANTHACEAE 421Subshrubs or perenni
- Page 55: ACANTHACEAE 423108. Strobilanthes d
- Page 58 and 59: 426ACANTHACEAESubshrubs 30-50 cm ta
- Page 60 and 61: 428ACANTHACEAEleaf blade lanceolate
- Page 62 and 63: 430ACANTHACEAEmm, apex gland-tipped
- Page 64 and 65: 432ACANTHACEAEusually attenuate and
- Page 66 and 67: 434ACANTHACEAEca. 4 cm, outside pub
- Page 68 and 69: 436ACANTHACEAEAbout 250 species: tr
- Page 70 and 71: 438ACANTHACEAEThis species is used
- Page 72 and 73: 440ACANTHACEAEEranthemum shweliense
- Page 74 and 75: 442ACANTHACEAEbrous with few sessil
- Page 76 and 77: 444ACANTHACEAE10b. Fertile bracts e
- Page 78 and 79: 446ACANTHACEAEbranous, abaxially gl
- Page 82 and 83: 450ACANTHACEAEJusticia betonica Lin
- Page 84 and 85: 452ACANTHACEAEside hispid, 4-lobed
- Page 86 and 87: 454ACANTHACEAE● On rocks, thicket
- Page 88 and 89: 456ACANTHACEAEcending, minutely whi
- Page 90 and 91: 458ACANTHACEAEmm, 5-lobed almost to
- Page 92 and 93: 460ACANTHACEAEovate, (2-)3-5.5 × 1
- Page 94 and 95: 462ACANTHACEAE1. Rhinacanthus beesi
- Page 96 and 97: 464ACANTHACEAEsometimes aggregated
- Page 98 and 99: 466ACANTHACEAEJusticia bivalvis Lin
- Page 100 and 101: 468ACANTHACEAE3. Hypoestes triflora
- Page 102 and 103: 470ACANTHACEAEbase muticous or spur
- Page 104 and 105: 472ACANTHACEAEin corolla tube; thec
- Page 106 and 107: 474ACANTHACEAEStamens 2, exserted f
- Page 108 and 109: 476ACANTHACEAE1a. Corolla outside p