16.01.2013 Views

Download - NERC Open Research Archive - Natural Environment ...

Download - NERC Open Research Archive - Natural Environment ...

Download - NERC Open Research Archive - Natural Environment ...

SHOW MORE
SHOW LESS

Create successful ePaper yourself

Turn your PDF publications into a flip-book with our unique Google optimized e-Paper software.

Somatochlora metallica (Vander Linden) Brilliant emerald<br />

Description S. metallica is slightly larger than the other two resident corduliid species, and<br />

has a greener appearance. It is readily distinguished from S. arctica and<br />

Cordulia aenea, which it resembles, by the distinctive pattern of yellow facial<br />

markings and the shape of the anal appendages. Also, in females, there is a<br />

very prominent spike-like vulvar scale which projects almost perpendicularly<br />

to the body on the ventral abdominal surface between segments 8 and 9, and<br />

is almost as long as those two segments. The eyes are green. The thorax is<br />

bright bronze-green and not so heavily downy as C. aenea. The abdomen,<br />

which is slightly club-shaped in males, is dark metallic green with a bronze<br />

sheen. There are pale yellow markings on abdominal segments 2 and 3, both<br />

laterally and ventrally and, in females, two small spots on segment 3 dorsally.<br />

There is a small amber area at the base of the wings which, in mature females,<br />

are entirely suffused with a pale amber tint.<br />

Habitat In south-east England, S. metallica breeds on mesotrophic, neutral to mildly<br />

acidic, ponds, lakes and canals, often where there are sheltered bays overhung<br />

with trees and bushes. It also occurs on small slow-flowing rivers. In the<br />

Scottish Highlands, S. metallica often breeds in similar habitat to C. aenea,<br />

namely: ancient peaty lochs which usually contain white water-lily, bogbean<br />

and marginal Sphagnum.<br />

Breeding biology On finding a female, copulation takes place away from water in trees or<br />

amongst low bushes and scrub. The female oviposits while hovering very low<br />

down, repeatedly tapping her abdomen, two to three times per second, into<br />

either wet Sphagnum moss or peat (both under and away from the water), or<br />

directly into shallow water amongst tree roots or common reed. Oviposition in<br />

S. metallica is discussed by Smith (1984) and Fox (1991). The larvae live<br />

amongst peaty detritus and leaf litter, and probably take two or more years to<br />

develop.<br />

Flight periods S. metallica is on the wing from mid-June (occasionally earlier) to mid-August.<br />

Status and distribution<br />

European and world<br />

distribution<br />

In Scotland it can be seen with C. aenea, Aeshna juncea,<br />

quadrimaculata and the commoner northern damselflies. In south-east<br />

England the list includes most of the species to be found on mesotrophic<br />

ponds and lakes. On slow-flowing rivers, it can be seen with Calopteryx<br />

splendens and Platycnemis penmpes.<br />

In Britain, S. metallica is divided into two distinct populations. One, in the<br />

Highlands, is centred on Glen Affric and Glen Mor. The other, in England, is<br />

much larger and includes the Weald of SusseX and west Kent, and the heaths of<br />

Surrey, north Hampshire and Berkshire. The first fully documented record of S.<br />

metallica in Britain was from Strathglass in 1869 by Dr Buchanan White<br />

(McLachlan 1870). At the turn of the century it was still thought to be solely a<br />

Scottish species by many leading entomologists (Lucas 1900), despite the fact<br />

that Stephens (1835-37) had recorded it as 'rare in England: it has been found<br />

in the metropolitan district in June'. It was first (re)discovered in England on 4<br />

August 1908 at The Warren, near Crowborough in Sussex, by E R Speyer<br />

(Speyer 1909). Considering that many of the entomologists of that period<br />

were based in south-east England, perhaps S. metallica was indeed absent,<br />

but one wonders whether, for example, the C. aenea recorded by C A Briggs<br />

on 13 September 1891 at Bookham, Surrey, was in fact S. metallica, in view of<br />

the exceptional late date. In 1922, S. metallica was found at Loch a' Chrion<br />

Diore in Argyllshire by K J Morton (specimen in the Royal Scottish Museum),<br />

and was refound there by E M and R W J Smith on 27 June 1995.<br />

S. metallica occurs in central and northern Europe and east to the Volga and<br />

Asia Minor. In the south of its range it is restricted to higher altitudes, such as<br />

the Pyrenees and the Alps.<br />

86

Hooray! Your file is uploaded and ready to be published.

Saved successfully!

Ooh no, something went wrong!