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Coenagrion armatum (Charpentier) Norfolk damselfly Description The male of C. armaturn is almost entirely black with dull greyish blue coloration on several segments at either extremity of the dorsal surface of the abdomen, though not segment 10 which is black. Unlike other species of this genus, there are no antehumeral stripes on the male thorax, although four spots may occur. The black marking on segment 2 of the abdomen typically has a lateral, sometimes isolated, arm extending along either side of the segment. The most notable feature of male C. armaturn, however, is the anal appendages, in which the inferior pair of appendages is very elongated. Females are mostly dull black with greenish markings, but they are unique in that the black on the dorsal surface of segment 2 does not extend the whole length of the segment — the anterior half is greenish. Of the female, Porritt (1910) wrote 'when flying in the sun, the mature insect is a very striking Agrion and cannot be mistaken for any other British species; the green on the front and posterior segments of the abdomen shines like emeralds...'. Habitat and C. armatum favours mesotrophic ponds and ditches with an abundance of breeding biology common reed and sedges, amongst which they fly Females have been observed to oviposit into the tissues of frogbit. Flight periods The flight period in Britain is not well recorded, but is likely to have been from Status and distribution European and world distribution late May to at least mid-July C. armaturn was first recorded in Britain at Stalham, Norfolk in May 1903 by Balfour-Browne (Balfour-Browne 1904). A Mr Edelston subsequently informed Balfour-Browne that he had found it there in 1902. The site, at Sutton Broad, Stalham, appears to have been the stronghold of the species, it being seen there regularly until the 1950s. Porritt (1912) recorded C. armatum 'on both Sutton and Stalham Broads, and over a fairly wide area'. A few C. armatum of both sexes were also recorded from a ditch at Hickling Broad by T A Coward on 24 May 1919. Cynthia Longfield (1954) wrote that A E Gardner had reported that C. armatum still occurred in fair numbers, but she went on to say that 'it is, however, very disturbing that the latter's restricted habitat is rapidly drying out and the nature of the vegetation changing completely.. and it is sincerely to be hoped that some remedy will be found in time to save the extinction of C. armatum'. It was not, and the species was last reported in 1957 (Hammond 1977). Surveys by the NCC in the mid-1970s found the area of its former site to be thoroughly overgrown with common reed, sallow and alder carr, and mostly dried up. C. armaturn is a boreal species which occurs from the Baltic area eastwards through Poland and Russia to Mongolia. It was last recorded in The Netherlands in 1924 (Geijskes & van Tol 1983), and was lost from sites in northern Germany in the mid-1970s owing to the drying up of its habitat (Schmidt 1978). 50
Coenagrion armatum (Charpentier) Norfolk damselfly 1975-90 o 1950-74 o Pre-195C 6 'EZ) 3 51 -(2 0
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Cover photograph: Aeshna juncea Ins
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@ NERC Copyright 1996 ISBN 0 11 701
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FOREWORD Having been on the fringe
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ACKNOWLEDGMENTS The following list
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Howard, G.; Howard, LW; Howe, M.; H
Page 13: Walker, D.S.; Walker, FA; WaLker, I
Page 16 and 17: CHECKLIST The following checklist i
Page 18 and 19: Aeshna mixta Latreille 1805 Aeschna
Page 20 and 21: THE BRITISH AND IRISH ODONATA IN A
Page 22 and 23: of distributions of dragonflies usi
Page 24 and 25: I 00 o o Figure 3. Map of coverage
Page 26 and 27: ,o " Figure 5. Map of coverage at t
Page 28 and 29: FIELDWORK AND DATA MANAGEMENT METHO
Page 30 and 31: areas, and encouraged in this by bo
Page 32 and 33: Table 3. Distribution of each speci
Page 34 and 35: Table 4 (con°. Vice-county distrib
Page 36 and 37: Table 4 (cont). Vice-county distrib
Page 38 and 39: Computerisation and validation Havi
Page 40 and 41: dominance of one individual over an
Page 42 and 43: RESIDENT SPECIES Description Habita
Page 44 and 45: Calopteryx splendens (Harris) Bande
Page 46 and 47: Description Flight periods Status a
Page 48: Description Lestes dryas Kirby Scar
Page 51 and 52: Platycnemis pennipes (Pallas) White
Page 53 and 54: Pyrrhosoma nymphula (Sulzer) 1975-9
Page 55 and 56: Erythromma najas (Hansemann) 1975-9
Page 57 and 58: Coenagrion mercuriale (Charpentier)
Page 59 and 60: Coenagrion scituluni (Rambur) Daint
Page 61 and 62: Coenagrion hastulatuni (Charpentier
Page 63: Coenagrion lunulatum (Charpentier)
Page 67 and 68: Coenagrion puella (Linnaeus) Azuxe
Page 69 and 70: Coenagrion pulchellum (Vander Linde
Page 71 and 72: Enallagma cyathigerurn (Charpentier
Page 73 and 74: Ischnura pumilio (Charpentier) Scar
Page 75 and 76: Ischnura elegans (Vander Linden) 19
Page 77 and 78: Ceriagrion tenellurn (Villers) 1975
Page 79 and 80: Aeshna caerulea (Strom) Azure hawke
Page 81 and 82: Aeshna juncea (Linnaeus) Common haw
Page 83 and 84: Montiaghs Moss, near Aghalee, Co An
Page 85 and 86: Sundon Springs chalk quarry, near L
Page 87 and 88: PLATE 1 Calopteryxvirgo adult PLATE
Page 89 and 90: PLATE 13 Libellula fulva adult PLAT
Page 91 and 92: Aeshna mixta Latreile Migrant hawke
Page 93 and 94: Aeshna cyanea (Muller) Southern haw
Page 95 and 96: Aeshna grandis (Linnaeus) Brown haw
Page 97 and 98: Aeshna isosceles (Muller) Norfolk h
Page 99 and 100: Anax imperator Leach Emperor dragon
Page 101 and 102: Brachytron pratense (Muller) Hairy
Page 103 and 104: Gomphus vulgatissimus (Linnaeus) 19
Page 105 and 106: Cordulegaster boltonii (Donovan) Go
Page 107 and 108: Cordulia aenea (Lin.naens) Downy em
Page 109 and 110: Somatochlora metallica (Vander Lind
Page 111 and 112: Somatochlora arctica (Zetterstedt)
Page 113 and 114: Oxygastra curtisii (Dale) Orange-sp
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Libellula quadrimaculata Linnaeus 1
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Libellula fulva Mailer Scarce chase
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Libellula depressa Linnaeus 1975-90
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Orthetrum cancellatum (Linnaeus) 19
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Orthetrum coerulescens (Fabricius)
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Sympetrum striolaturn (Charpentier)
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Sympetrum sanguineum (Muller) Ruddy
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Sympetruni danae (Sulzer) 1975-90 o
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Leuconhinia dubia (Vander Linden) W
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IMIVLIGRANTS AND ACCIDENTAL SPECIES
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POSSIBLE ADDITIONS TO THE BRITISH A
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SEASONAL OCCURRENCE PREVIOUS REPRES
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700 600 500 400 300. 200 100 0 90 8
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70 60 50 40 30 20 10 90 80 70 60 50
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Finally, differences in recorder be
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ODONATA LOCALITY 6411 V10E-COUNTY N
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Particularly in the case of residen
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conditions and which may be of grea
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1960s from its single locality alon
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pond. One of the authors did this o
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REFERENCES Aguilar, J. d', Dommange
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Gabb, R. & Kitching, D. 1992. The d
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Coenagrion rnercuriale (Charpentier
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Dee'. Transactions of the Natural H
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Circumboreal Occurring in a band en
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Marl A whitish clay soil with a hig
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Vulvar scale WATCH Weald Wheel posi
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Appendix 2. LIST OF PLANT NAMES Pla
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Libefiula depressa 5,6,14,17,18,19,
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