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Status and distribution<br />
Aeshna grandis (Linnaeus) Brown hawker<br />
Description Both sexes of this species are unmistakable in having a dark brown body with<br />
wings suffused all over with an orange-brown tint. The sides of the thorax<br />
possess two broad yellow bands. In males, there are a pair of large blue spots<br />
on the dorsal surface of abdominal segment 2, a large blue mark laterally on<br />
segment 3, and smaller blue marks along the sides of segments 4-8. There<br />
are also tiny paired yellow markings dorsally on abdominal segments 2-8. The<br />
eyes are brownish blue. In females, the blue spots on segment 2 are absent,<br />
and the marks along the sides of the abdomen are yellow<br />
Habitat A. grandis occurs on ponds, lakes, slow-moving rivers, canals and ditches. It is<br />
one of the most frequently encountered large dragonflies in south-eastern and<br />
midland counties of Britain, often visiting garden ponds and flying late into the<br />
evening.<br />
Breeding biology A. grandis is territorial and adult male densities rarely exceed 2 per 100 m of<br />
water's edge. It is unusual also in defending territories away from water. After<br />
a lengthy copulation, the female lays her eggs singly into the tissues of<br />
emergent plants just below the water surface, or into floating aquatic<br />
vegetation provided there is sufficient support for her not to become<br />
completely immersed. Sometimes several females will settle on a small<br />
floating log and oviposit close together into the wood, whilst at other times<br />
they will show aggressive behaviour towards each other. The eggs pass the<br />
winter in diapause. The larvae take from two to four years to complete<br />
development, and live amongst submerged water plants. As with several other<br />
weed-dwelling aeshnid species, such asil. cyanea and Anax imperator, a<br />
proportion of first-year larvae are cryptically coloured, often with a banded<br />
pattern. This may help reduce cannibalism and interspecific predation from<br />
older larvae.<br />
Flight periods A. grandis is on the wing from early July to early October. It occurs with the<br />
many other species of dragonfly which breed in well-vegetated ponds and<br />
waterways within its geographical range.<br />
In Britain, A. grandis is widespread throughout the lowland areas of the southeast,<br />
the midlands and as far north as Lancashire, Yorkshire, and the southern<br />
fringe of Cumbria, with a few isolated records further north. There is a single<br />
record from Scotland: from Colvend in Dumfries-shire in the mid-19th century.<br />
A. grandis occurs in Wales along the arm of the Montgomery canal, and near<br />
the border with Cheshire. It is absent from Devon and Cornwall. In Ireland, A.<br />
grandis is probably more common in the central midlands and the south-east<br />
than the records suggest — most of the fieldwork there having been done in the<br />
early summer — but it does appear to be rare in some western districts,<br />
notably western Co Cork and Co Kerry.<br />
European and world On the continent A. grandis is found from France to eastern Siberia. It occurs at<br />
distribution quite high latitudes in Scandinavia, but is absent from much of the<br />
Mediterranean area.<br />
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