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Status and distribution<br />

Aeshna grandis (Linnaeus) Brown hawker<br />

Description Both sexes of this species are unmistakable in having a dark brown body with<br />

wings suffused all over with an orange-brown tint. The sides of the thorax<br />

possess two broad yellow bands. In males, there are a pair of large blue spots<br />

on the dorsal surface of abdominal segment 2, a large blue mark laterally on<br />

segment 3, and smaller blue marks along the sides of segments 4-8. There<br />

are also tiny paired yellow markings dorsally on abdominal segments 2-8. The<br />

eyes are brownish blue. In females, the blue spots on segment 2 are absent,<br />

and the marks along the sides of the abdomen are yellow<br />

Habitat A. grandis occurs on ponds, lakes, slow-moving rivers, canals and ditches. It is<br />

one of the most frequently encountered large dragonflies in south-eastern and<br />

midland counties of Britain, often visiting garden ponds and flying late into the<br />

evening.<br />

Breeding biology A. grandis is territorial and adult male densities rarely exceed 2 per 100 m of<br />

water's edge. It is unusual also in defending territories away from water. After<br />

a lengthy copulation, the female lays her eggs singly into the tissues of<br />

emergent plants just below the water surface, or into floating aquatic<br />

vegetation provided there is sufficient support for her not to become<br />

completely immersed. Sometimes several females will settle on a small<br />

floating log and oviposit close together into the wood, whilst at other times<br />

they will show aggressive behaviour towards each other. The eggs pass the<br />

winter in diapause. The larvae take from two to four years to complete<br />

development, and live amongst submerged water plants. As with several other<br />

weed-dwelling aeshnid species, such asil. cyanea and Anax imperator, a<br />

proportion of first-year larvae are cryptically coloured, often with a banded<br />

pattern. This may help reduce cannibalism and interspecific predation from<br />

older larvae.<br />

Flight periods A. grandis is on the wing from early July to early October. It occurs with the<br />

many other species of dragonfly which breed in well-vegetated ponds and<br />

waterways within its geographical range.<br />

In Britain, A. grandis is widespread throughout the lowland areas of the southeast,<br />

the midlands and as far north as Lancashire, Yorkshire, and the southern<br />

fringe of Cumbria, with a few isolated records further north. There is a single<br />

record from Scotland: from Colvend in Dumfries-shire in the mid-19th century.<br />

A. grandis occurs in Wales along the arm of the Montgomery canal, and near<br />

the border with Cheshire. It is absent from Devon and Cornwall. In Ireland, A.<br />

grandis is probably more common in the central midlands and the south-east<br />

than the records suggest — most of the fieldwork there having been done in the<br />

early summer — but it does appear to be rare in some western districts,<br />

notably western Co Cork and Co Kerry.<br />

European and world On the continent A. grandis is found from France to eastern Siberia. It occurs at<br />

distribution quite high latitudes in Scandinavia, but is absent from much of the<br />

Mediterranean area.<br />

72

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