Download - NERC Open Research Archive - Natural Environment ...
Download - NERC Open Research Archive - Natural Environment ...
Download - NERC Open Research Archive - Natural Environment ...
Create successful ePaper yourself
Turn your PDF publications into a flip-book with our unique Google optimized e-Paper software.
accessary genitalia of males and the prominent<br />
vulvar scale of females are diagnostic. S. vulgatum<br />
breeds in ditches, ponds, lakes and slow-flowing<br />
rivers, and is on the wing from early July to<br />
October. It has been recorded in Britain on fewer<br />
than ten occasions, mostly from the London area,<br />
but also once from Torquay, Devon, and once from<br />
Kingston upon Hull, Yorkshire. None of the records<br />
is from recent years. It has been recorded several<br />
times from Jersey (Le Quesne 1946), but not from<br />
Ireland. On the continent, S. vulgatum occurs from<br />
eastern France to southern Scandinavia, and east to<br />
China. It appears to have increased in The<br />
Netherlands in recent years (Geijskes & van Tol<br />
1983), and so may reach Britain more frequently in<br />
the future.<br />
Sympetrum fonscolombii (Sélys-Longchamps)<br />
In mature male S. fonscolombil, the abdomen is<br />
brighter red than S. stholatum and more<br />
parallel-sided. The veins in the proximal halves of<br />
the wings are red, thus distinguishing S.<br />
fonscolombii from all other Sympetrum species. In<br />
females these veins are yellowish and the abdomen<br />
is yellow-brown. In both sexes the pterostigmata are<br />
pale with a conspicuous black border along two<br />
sides, and there is a saffron patch at the base of the<br />
wings which may be quite extensive, though never<br />
as much as in S. flaveolum. It breeds in shallow static<br />
water bodies, and is on the wing from mid-June to<br />
October, earlier in southern Europe, where it may<br />
produce two generations in one year. S.<br />
fonscolombii is migratory and occurs sporadically in<br />
Britain, usually in mid- to late summer, although a<br />
male was recorded on 9 May 1987 at Broornhill<br />
Burrows, Pembrokeshire. The majority of records<br />
are from southern England, but it has occurred as<br />
far north as the Isle of May on the east Scotland<br />
coast, on 17 August 1911. There are two records<br />
from Co Cavan, Ireland, on 20 July 1941 and 27 July<br />
1942. It has almost certainly bred occasionally in<br />
southern England (Longfield 1949c), but has been<br />
unable to sustain a population. It has bred on Jersey,<br />
where Le Quesne (1946) noted oviposition taking<br />
place in a canal on 11 May 1945. He continued to<br />
see adults there until July, then in August he found an<br />
immature female, the first specimen in this condition<br />
that he had seen that year. On 30 August, many<br />
exuviae and teneral adults were located at the site.<br />
He suggested that the evidence could indicate that<br />
the species may overwinter in the adult state. It is<br />
more probable that S. fonscolombii had undergone<br />
two generations in one year, which this species is<br />
known to,accomplish in more southern latitudes,<br />
and that Le Quesne had missed a synchronous<br />
spring emergence toward the end of April. On the<br />
continent, S. fonscolombii is widespread in the<br />
Mediterranean region, and is found east to India,<br />
Mongolia and the Pacific. It occurs throughout<br />
Africa.<br />
112<br />
Sympetrum flaveolum (Linnaeus)<br />
In S. flaveolum the parallel-sided abdomen is red in<br />
mature males, yellowish brown in females. The<br />
species is distinctive for having a broad saffronyellow<br />
patch on the basal area of each wing. This is<br />
much more extensive than on any other Sympetrum<br />
except, occasionally, S. fonscolombii from which it<br />
can be distinguished by its black wing venation and<br />
dark pterostigma. It breeds in marshy ponds,<br />
ditches, and lakes, and the backwaters of slowflowing<br />
rivers. It is on the wing from late June to<br />
October. S. flaveolum is migratory and occurs<br />
sporadically in Britain, mainly in the south although<br />
it occurred as far north as Keiss, Caithness, on 18<br />
July 1945. It appears to be recorded less frequently<br />
now than it was before 1950. There is strong<br />
circumstantial evidence to suggest that S. flaveolum<br />
has bred in southern England on a few occasions<br />
(Longfield 1949c), but has been unable to sustain a<br />
population. It has been recorded from Jersey, and<br />
Belle (1980) suggests that it bred formerly on<br />
Guernsey and Alderney. There are no records from<br />
Ireland. It is found throughout Europe except the<br />
extreme south and north, and occurs east to Siberia<br />
and Japan. It is absent from Africa.<br />
Pantala flavescens (Fabricius)<br />
This brown libellulid is larger than any of the<br />
resident libellulids in Britain or Ireland, having an<br />
average overall length of 49-52 mrn. The abdomen<br />
of both sexes is yellow-brown with a black<br />
longitudinal line, of uneven width, along the middorsal<br />
surface which thickens posteriorly into black<br />
triangular marks on seyments 8-10. The thorax is<br />
brown dorsally with yellow-brown sides. The wings<br />
of males may develop a small pale brown patch<br />
apically. There are two records from Britain. One is<br />
of a specimen captured in 1823 at Horning, Norfolk,<br />
by J Sparshall. This record is discussed by Fraser<br />
(1956) who had experience of it in India, and who,<br />
despite the doubts expressed by some leading<br />
entomologists of the 19th century as to how P<br />
flavescens could have reached Britain, preferred to<br />
keep an open mind. The second record is of an<br />
individual taken at Bolton, Lancashire, in July 1951<br />
by A Hazelwood, and which is now in the National<br />
Museums on Merseyside, Liverpool. It is believed<br />
to have arrived in Britain as a ship-borne immigrant<br />
with a consignment of bananas. Longfield (in<br />
Corbet et al, 1960) cites a case of a male P<br />
flavescens sent to her by 0 G Watkins of Plymouth<br />
which was found flying around the wardroom of a<br />
British warship a few days before reaching<br />
Devonport Harbour from Singapore in 1955. It is a<br />
pantropical species, capable of migrating vast<br />
distances. It is found in Asia, Australasia, the<br />
Americas, Africa and many of the islands in the<br />
Indian Ocean and the Pacific Ocean. It only occurs<br />
sporadically in Europe.