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accessary genitalia of males and the prominent<br />

vulvar scale of females are diagnostic. S. vulgatum<br />

breeds in ditches, ponds, lakes and slow-flowing<br />

rivers, and is on the wing from early July to<br />

October. It has been recorded in Britain on fewer<br />

than ten occasions, mostly from the London area,<br />

but also once from Torquay, Devon, and once from<br />

Kingston upon Hull, Yorkshire. None of the records<br />

is from recent years. It has been recorded several<br />

times from Jersey (Le Quesne 1946), but not from<br />

Ireland. On the continent, S. vulgatum occurs from<br />

eastern France to southern Scandinavia, and east to<br />

China. It appears to have increased in The<br />

Netherlands in recent years (Geijskes & van Tol<br />

1983), and so may reach Britain more frequently in<br />

the future.<br />

Sympetrum fonscolombii (Sélys-Longchamps)<br />

In mature male S. fonscolombil, the abdomen is<br />

brighter red than S. stholatum and more<br />

parallel-sided. The veins in the proximal halves of<br />

the wings are red, thus distinguishing S.<br />

fonscolombii from all other Sympetrum species. In<br />

females these veins are yellowish and the abdomen<br />

is yellow-brown. In both sexes the pterostigmata are<br />

pale with a conspicuous black border along two<br />

sides, and there is a saffron patch at the base of the<br />

wings which may be quite extensive, though never<br />

as much as in S. flaveolum. It breeds in shallow static<br />

water bodies, and is on the wing from mid-June to<br />

October, earlier in southern Europe, where it may<br />

produce two generations in one year. S.<br />

fonscolombii is migratory and occurs sporadically in<br />

Britain, usually in mid- to late summer, although a<br />

male was recorded on 9 May 1987 at Broornhill<br />

Burrows, Pembrokeshire. The majority of records<br />

are from southern England, but it has occurred as<br />

far north as the Isle of May on the east Scotland<br />

coast, on 17 August 1911. There are two records<br />

from Co Cavan, Ireland, on 20 July 1941 and 27 July<br />

1942. It has almost certainly bred occasionally in<br />

southern England (Longfield 1949c), but has been<br />

unable to sustain a population. It has bred on Jersey,<br />

where Le Quesne (1946) noted oviposition taking<br />

place in a canal on 11 May 1945. He continued to<br />

see adults there until July, then in August he found an<br />

immature female, the first specimen in this condition<br />

that he had seen that year. On 30 August, many<br />

exuviae and teneral adults were located at the site.<br />

He suggested that the evidence could indicate that<br />

the species may overwinter in the adult state. It is<br />

more probable that S. fonscolombii had undergone<br />

two generations in one year, which this species is<br />

known to,accomplish in more southern latitudes,<br />

and that Le Quesne had missed a synchronous<br />

spring emergence toward the end of April. On the<br />

continent, S. fonscolombii is widespread in the<br />

Mediterranean region, and is found east to India,<br />

Mongolia and the Pacific. It occurs throughout<br />

Africa.<br />

112<br />

Sympetrum flaveolum (Linnaeus)<br />

In S. flaveolum the parallel-sided abdomen is red in<br />

mature males, yellowish brown in females. The<br />

species is distinctive for having a broad saffronyellow<br />

patch on the basal area of each wing. This is<br />

much more extensive than on any other Sympetrum<br />

except, occasionally, S. fonscolombii from which it<br />

can be distinguished by its black wing venation and<br />

dark pterostigma. It breeds in marshy ponds,<br />

ditches, and lakes, and the backwaters of slowflowing<br />

rivers. It is on the wing from late June to<br />

October. S. flaveolum is migratory and occurs<br />

sporadically in Britain, mainly in the south although<br />

it occurred as far north as Keiss, Caithness, on 18<br />

July 1945. It appears to be recorded less frequently<br />

now than it was before 1950. There is strong<br />

circumstantial evidence to suggest that S. flaveolum<br />

has bred in southern England on a few occasions<br />

(Longfield 1949c), but has been unable to sustain a<br />

population. It has been recorded from Jersey, and<br />

Belle (1980) suggests that it bred formerly on<br />

Guernsey and Alderney. There are no records from<br />

Ireland. It is found throughout Europe except the<br />

extreme south and north, and occurs east to Siberia<br />

and Japan. It is absent from Africa.<br />

Pantala flavescens (Fabricius)<br />

This brown libellulid is larger than any of the<br />

resident libellulids in Britain or Ireland, having an<br />

average overall length of 49-52 mrn. The abdomen<br />

of both sexes is yellow-brown with a black<br />

longitudinal line, of uneven width, along the middorsal<br />

surface which thickens posteriorly into black<br />

triangular marks on seyments 8-10. The thorax is<br />

brown dorsally with yellow-brown sides. The wings<br />

of males may develop a small pale brown patch<br />

apically. There are two records from Britain. One is<br />

of a specimen captured in 1823 at Horning, Norfolk,<br />

by J Sparshall. This record is discussed by Fraser<br />

(1956) who had experience of it in India, and who,<br />

despite the doubts expressed by some leading<br />

entomologists of the 19th century as to how P<br />

flavescens could have reached Britain, preferred to<br />

keep an open mind. The second record is of an<br />

individual taken at Bolton, Lancashire, in July 1951<br />

by A Hazelwood, and which is now in the National<br />

Museums on Merseyside, Liverpool. It is believed<br />

to have arrived in Britain as a ship-borne immigrant<br />

with a consignment of bananas. Longfield (in<br />

Corbet et al, 1960) cites a case of a male P<br />

flavescens sent to her by 0 G Watkins of Plymouth<br />

which was found flying around the wardroom of a<br />

British warship a few days before reaching<br />

Devonport Harbour from Singapore in 1955. It is a<br />

pantropical species, capable of migrating vast<br />

distances. It is found in Asia, Australasia, the<br />

Americas, Africa and many of the islands in the<br />

Indian Ocean and the Pacific Ocean. It only occurs<br />

sporadically in Europe.

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