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The larval ecology of most species is poorly known,<br />

but it is apparent that only a small part of a wetland<br />

complex may provide suitable larval habitats, even if<br />

the adults range widely over the whole site, as is<br />

often the case with the larger Anisoptera. Proof of<br />

breeding is often crucial in site evaluation and<br />

protection, and larvae and exuviae provide the best<br />

possible evidence.<br />

Finally, the larvae of Odonata are, in a very different<br />

way, as striking as the adults, often subtly coloured<br />

and camouflaged, and structurally diverse. In an<br />

Order containing rather few British and Irish<br />

species, the larvae can provide an extra challenge,<br />

requiring a new set of field skills for the observer<br />

who has already mastered the adults.<br />

Transect methods<br />

As early as the 1950s, the counting of adult male<br />

Odonata along a waterside transect was being used<br />

to monitor dragonfly populations (Moore 1953,<br />

1964, 1991c). The method has since become more<br />

widely known through its development for butterfly<br />

monitoring (Moore 1975; Pollard 1979, 1992).<br />

Butterflies are perhaps even better suited to transect<br />

counts than Odonata: most butterflies are not<br />

strongly territorial, and unlike many Odonata do not<br />

move far from their breeding sites. However, if<br />

counts of adult male Odonata are repeated regularly<br />

(ideally, once a week) in suitable weather conditions<br />

(sunny and with little wind) and close to mid-day,<br />

they can provide results which are comparable and<br />

thus a simple annual index of abundance. This<br />

method is very useful in quantifying flight period<br />

(see below), and can detect long-term trends in a<br />

species, perhaps reflecting site management<br />

(Brooks 1993); Moore (1991c) follows the pattern of<br />

colonisation and faunal change at ponds at<br />

Woodwalton Fen over 27 years using transect<br />

methods.<br />

Comprehensive monitoring<br />

Useful though transect counts can be, they are<br />

probably best used as part of a wider-ranging<br />

programme of monitoring. A review of the<br />

techniques for comprehensive monitoring has been<br />

provided by Moore and Corbet (1990). They<br />

discuss the need for monitoring, and aspects of the<br />

behaviour of Odonata which influence the ways in<br />

which methods may be applied. They conclude that<br />

the choice of survey methods will depend on the<br />

habitats involved, especially the accessibility of the<br />

water's edge, and the time available for monitoring.<br />

They make several recommendations.<br />

The preferred technique for monitoring<br />

Anisoptera is by counting exuviae, which should<br />

be collected as frequently as possible,<br />

throughout the emergence period.<br />

15<br />

Because of the difficulty of finding and<br />

identifying their exuviae, Zygoptera are best<br />

monitored by counting teneral adults, as<br />

frequently as possible, throughout the<br />

emergence period.<br />

Adult counts are comparable only when<br />

restricted to adult males by water, and when<br />

made within 1-2 hours of mid-day on sunny days<br />

with little or no wind.<br />

Whatever methods of assessment are used, it is<br />

preferable if they can be applied at regular<br />

intervals, and at the same time each day.<br />

As a minimum, Moore and Corbet recommend that<br />

counts of anisopteran exuviae, zygopteran teneral<br />

adults, and mature males by water should be made<br />

once a week throughout the season. With this<br />

required level of commitment, it is important to<br />

establish the purpose of monitoring, and to decide<br />

what level of detail is required. Can monitoring be<br />

focused on one or two species of particular<br />

importance? If management is being monitored, can<br />

representative sections of habitat be chosen rather<br />

than attempting to monitor a whole site? Does the<br />

monitoring need to be repeated annually? (A<br />

thorough survey every third year may be more -<br />

useful than incomplete or inconsistent surveys every<br />

year.)<br />

DESCRIPTION OF DATA SET<br />

The maps presented in this Atlas are based on all<br />

records received by the Odonata Recording<br />

Scheme up to the end of 1990. Some later records,<br />

of particular importance, are mentioned in the<br />

species accounts and elsewhere, but do not appear<br />

on the maps.<br />

Recording is a continuing process, because species<br />

distributions are not static and there remain large<br />

gaps in our knowledge of the biology of even the<br />

commonest species. The methods of data collection<br />

and validation described in this section refer both to<br />

the data included in this Atlas and to ongoing<br />

recording.<br />

Management of data collection<br />

The history of the Scheme and data collection has<br />

already been described. For most of the duration of<br />

the Recording Scheme, a national scheme organiser<br />

has been responsible for collating incoming records<br />

and providing feedback to recorders. This feedback<br />

has included comments on the significance of the<br />

records supplied, and requests for further<br />

information in support of particularly outstanding<br />

records. Additionally, through the newsletters,<br />

recorders have been urged to visit under-worked

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