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48 U.S. NATIONAL MUSEUM BULLETIN 25 7 The characteristic genus, recognized and resurrected by Obraztsov (personal communication) and by Hannemann (1961), contains in the Palaearctic and Palaeotropic regions several very similar species, hitherto mostly attributed to ^'Eudemis" purpurissatana Kennel, which species, however, is not even congeneric, as ascertained by Falkovitch recently. It belongs to Eudemopsis Falkovitch, 1962. The genus Eudemis represents a large complex of species which I propose to divide into two subgenera. The subgenus Eudemis Hiibner, new status, comprises chiefly Palaearctic species, of which the males have no additional coremata and posses a simple valva. The subgenus Acanthothyspoda Lower, new status, contains tropical species, usually greenish-colored, males \vith. abundantly hau-ed posterior tibiae, with lateral pairs of hair pencils, being additional coremata on most abdominal segments, with a more complicated valva, and often with socii spiny at the top. The peculiar external characters are restricted to the males and should, consequently, be regarded as secondary sexual characters only. They are absent in true Eudemis but not well developed in some tropical Acanthothyspoda. The specialized (plesiomorph) Eudemis species and apomorph Acanthothyspoda species appear to approach each other closely, linked by intermediate forms. Therefore, there is, in my opinion, no reason for generic separation of the two groups. The females of both are simOar. In Clarke's monograph (1958, vol. 3) not less than 11 species, recorded under Olethreutes, are illustrated (see pp. 49, 51, 52). Photographs of the females of two species have been reproduced by Clarke. The genus may be redescribed as foUows: Species of moderate size, usually with a big round terminal spot in the forewing, tropical species more or less green-tinged. Head with loosely appressed scales, roughish on crown. Proboscis developed. Ocellus posterior. Antenna in male thickened, fasciculateciliated, cUiations 1/2. Palpus moderately long, porrected, median segment strongly dilated, triangular, terminal segment moderate, exposed, slightly drooping, obtuse. Thorax with a slight double posterior crest. Posterior tibia in male sometimes extremely dilated and expanded by pencils, whorls or brushes of dense hairs of diverse shape. Abdomen in male often normally haired and scaled, but sometimes with a pair of long hair-pencils at the sides of each segment; these pencils are implanted on segments 2-3, sometimes on more segments, on distinct projecting papillae, or on raised oval fields. Forewing elongate-subtruncate, moderately broad, termen usually slightly rounded, little oblique. Vein 2 from beyond 2/3, 3 from angle, close to or rather distant from 4, 5 distant, 6 slightly diverging from 5 towards base, 7 separate, to termen, 7 to 9 equidistant at base, seldom 7 and 8 almost connate, 11 from before middle of cell,
MICROLEPIDOPTERA OF PHILIPPINE ISLANDS 49 upper parting vein developed, from just bejônd 10 or from about 2/3 between 11 and 10 to between 6 and 7; lower parting vein also developed, sinuate, from towards base to below base of 4. Hindwing wîth a cubital pecten; with tornus sometimes slightly modified, covered with modified scales, sometimes causing anomal venation. Consequently, this venation is subject to considerable variation in the male. Vein 2 from 1/2-2/3, 3 from before angle, 4 remote from angle, or 3 and 4 connate from angle, 5 approximated at base, 6 and 7 closely approximated toward base. Type species: Subgenus Eudeinis Hiibner, 1826, new status Tortrix porphyranaHubner. Male genitalia: Tegumen elongate, not conspicuously narrowed towards top. Valva usually slender, curved (seldom dUated and rounded), sacculus indefinite; Spci, Spc2, and sometimes Spcg may all be present. Gnathos pending, arms long, strongly bristled or spined at the top. Female genitalia with a short, funnel-shaped sterigma, and a short, cyUndrical coUiculum. Signa, two flattened horns. The following species should be transferred to the present genus. Argyroploce archimedias Meyrick, 1912 (China); Clarke, 1958, vol. 3, p. 540, pi. 269, figs. 4-4a. Eucosma mniochlora Meyrick, 1916 (India, China); Clarke, 1958, vol. 3, p. 528, pi. 263, figs. 4â. Argyroploce pompholycias Meyrick, 1935 (China); Clarke, 1958. vol. 3, p. 539, pi. 268, figs. 2-2a. Key to the PhUippiae Species of Subgenus Eudemis 1. Forewing with the dorsal spot reaching above middle of wing, its top pointing to costa before anterior edge of transverse fascia . . . . E. archimedias Forewing with the dorsal spot reaching to middle of wing, its top directed exactly to anterior edge of transverse fascia on costa . . . . E. oxygona Eudemis (Eudemis) archim.edias (Meyrick, 1912), new combination Figure 526 Argyroploce archimedias Meyrick, 1912, Exotic Microlepidoptera, vol. 1, p. 63 (cf, China).— Clarke, 1955, Catalogue . . . Microlepidoptera . . . Meyrick, vol. 1, p. 52. Oiei/ireu^fspurpi/msa/a/m Clarke, 1958 (not Kennel, 1901), Catalogue . . . Microlepidoptera . . . Meyrick,vol.3, p. 540, pi. 269, figs. 4-4a (arc/iimedtas Meyrick, syn.).—Falkovitch, 1962, Rev. Ent. USSR, vol. 41, p. 191 (archimedias Meyrick, a distinct species). It may be stipulated th&i " Penthiiia" purpurissatana Kennel, 1901 does not belong to the present group of species, in spite of the very
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Appendix 1 Selection of Lectotypes
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Appendix 2 List of Selected Lectoty
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Figures 370-373.—Male genitalia o
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Index of Latin names of Food-plants
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