Appendix F Detailed Cover Type Tables - USDA Rural Development
Appendix F Detailed Cover Type Tables - USDA Rural Development
Appendix F Detailed Cover Type Tables - USDA Rural Development
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Bemidji – Grand Rapids<br />
Biological Assessment and Evaluation<br />
4.2.5.4 Triangle Moonwort (Botrychium lanceolatum var. angustisegmentum)<br />
Botrycium lanceolatum var. angustisegmentum is listed as DNR Threatened, CNF RFSS and DRM<br />
Threatened.<br />
Species Description<br />
Botrychium lanceolatum has a stalk 6‐15 cm (2.4‐5.9 in.) long, with an unstalked frond. The frond (sterile<br />
blade) is dark green, very shinny, glabrous, and deltoid. It has two to five pairs of pinnatifid pinnae, the<br />
lowest pair much the larger (hence the common name triangle). The bud is enclosed in the stalk base.<br />
B. lanceolatum can be distinguished from B. matricariifolium (matricary grapefern) by the deltoid,<br />
usually sessile, blade with both sterile and fertile blades reflexed in bud. The variety of B. lanceolatum<br />
(S.G. Gmelin) in Minnesota is var. angustisegmentum. (Pease & Moore).<br />
Leaves of this species emerge in the spring and are divided into a sterile photosynthetic portion and a<br />
fertile spore‐bearing portion. Spores of the fertile portion mature slowly through the summer, changing<br />
to a noticeable gold color by late summer. The aboveground portion of the plant is killed by frost in the<br />
fall, sometimes as late as October.<br />
Species Habitat<br />
According to the DNR Rare Species Guide:<br />
In Minnesota, B. lanceolatum prefers moist, shady, mature northern hardwood forests, particularly in<br />
low areas. It usually occurs with Acer saccharum (sugar maple), Betula alleghaniensis (yellow birch),<br />
Fraxinus pennsylvanica (green ash), Fraxinus nigra (black ash), Quercus rubra (red oak), Tilia americana<br />
(basswood), and sometimes Thuja occidentalis (northern white cedar). The understory can be rather<br />
open and the ground cover often sparse with Athyrium angustum (lady fern) and other Botrychium<br />
species, especially B. matricariaefolium. Botrychium lanceolatum populations often occur as a few<br />
scattered individuals, but occasionally 50 or more can be found in a single site.<br />
Species Distribution and Occurrences within the Study Area<br />
Risk Factors<br />
A widely distributed plant, B. lanceolatum occurs from Newfoundland<br />
across the Canadian provinces to Yukon and Alaska, south to<br />
California, east to New Mexico, then Minnesota to the Atlantic Coast<br />
states as far south as Virginia and inland to Tennessee. The species is<br />
not noted in the central plains states west and south of Minnesota<br />
(DNR, 2009). <strong>USDA</strong> Plants Database at:<br />
http://plants.usda.gov/java/county?state_name=Minnesota&statefips<br />
=27&symbol=BOLA<br />
Botrychium lanceolatum is listed by the DNR and CNF with point data<br />
indicating 10 distinct populations identified throughout the Study<br />
Area.<br />
Threats to Botrychium lanceolatum include road building, timber harvesting, hydrologic alteration,<br />
invasion by exotic plant species, removal of mature hardwood forests, and exotic earthworm invasion.<br />
Invasion of exotic earthworms appears to represent the greatest threat to this species, and the genus as<br />
a whole within the CNF and LLR area. Herbivory and drought likely pose short term risks to this species.<br />
The single most important limiting factor affecting Botrychium sp. establishment, distribution and<br />
abuncance, according to Johnson‐Groh (1999 cf. Chadde and Kudray, 2001) is the relationship that this<br />
July 2010 Species and Associated Habitat Page 4‐46