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42 =>
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Table of Contents Declaration ii Ac
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4 Laboratory investigation into mic
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temperatures on fluorescence parame
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always been real sweeties. Also tha
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Chl Chlorophyll Chl a Chlorophyll a
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CHAPTER ONE GENERAL INTRODUCTION Wo
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(, 'I, I 1)CI ii I lU RH araphid (n
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dpl. II (I c I) cIq 111 I-Itro-ducJ
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c 11 cIa 11 (T-oduct-Ioll integrate
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( lapRl Wilclul IfInOdik 11011 impo
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Chdpt(1111 1. -, -- I--, -- -- I, -
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(-Jlaptýýr j( rcllcj-ý11 ý: - o
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1.3.2. The Photosynthetic carbon re
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( CflCI iii I TI )dU( t Zingmark 19
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( lidptLI ( ft fln JI flU UII Di wh
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Cha ter I (iclicl-al IntroiftwOoll
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whilst cells minimise their exposur
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1. Pigments Introduction Chlorophyl
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Spectral reflectance studies examin
- Page 41 and 42: (-'Ilaptcýr J, I, -, --, --, --, -
- Page 43 and 44: ( anci JII )tI( )(Jtk IO) which buf
- Page 45 and 46: ( ! p1'1 -- -1 -ý T-( ý 1)era J-1
- Page 47 and 48: Uh Temporal changes in the microsca
- Page 49 and 50: (I (rcI IpuiE't, cells as they emer
- Page 51 and 52: ( hap'wv -1 ( ýI II ntrodt 1ý t I
- Page 53 and 54: ( hapi"T 1 11 1- 11 1- ---- -- -- 1
- Page 55 and 56: (A lal )te II- iý; 11 11 1 1- IIIJ
- Page 57 and 58: ( 1ý lv-ý ý12, -) -, - -- - -- -
- Page 59 and 60: 2.1.3. The Colne Estuary "VIt"utc-1
- Page 61 and 62: (I --- -- "I'l- I -11-1--l- -I--"--
- Page 63 and 64: E I I Uneven surface topography ma]
- Page 65 and 66: 2.4. Wet bulk density (Equation 2.2
- Page 67 and 68: ('liapti 2- 2.5.4. Cell identificat
- Page 69 and 70: ( hapt )- acetone and DMF do not di
- Page 71 and 72: Chapter 2 Materials and Nlethocts P
- Page 73 and 74: Prior to a run all solvents were ge
- Page 75 and 76: (h1ft 'ý, I (I ', tý7! ýI)iI"Iý
- Page 77 and 78: QLapteLi2 2.6.6. Troubleshooting th
- Page 79 and 80: ---- ------ at 486.5 nm on a Cecil
- Page 81 and 82: Figure 2.10. A) The fibre optic mea
- Page 83 and 84: tcl -1 Materials jnd 'Mclho(k attac
- Page 85 and 86: Chapt r2 Materials mid Methods A hi
- Page 87 and 88: 2 2.9.2. Fluorescence parameters Fl
- Page 89 and 90: CHAPTER THREE ! i()fi1I1I 1]lILli\L
- Page 91: - -I ii-ofj) II 1- 1-() 111 llatim
- Page 95 and 96: tcr-3- I- --- I-, -- - --, -- -- --
- Page 97 and 98: tu -0 Testing the model of Underwoo
- Page 99 and 100: Biof-11111 lol-Illatioll Lind dc\ c
- Page 101 and 102: over days 3-14 appeared linear and
- Page 103 and 104: (_ hpii 3.3.2. Laboratory Investiga
- Page 105 and 106: -(- II Water content mi-Ijill Imill
- Page 107 and 108: _F 120 100 80 60 40 20 160- 140- 12
- Page 109 and 110: 0) '7 CY) 6- E =- 1000 0 ci) 0 0246
- Page 111 and 112: _0 2.0 1.5 1.0 0.5 0.0 Time (days)
- Page 113 and 114: R, -jOiln 1,1 11ý"; - I, d dlýý\
- Page 115 and 116: cq 4.0 3.5 3.0 E 2.5 z -0 U) 2.0 1.
- Page 117 and 118: E 4.0- 3.5- 3.0- 2.5- 2.0- 1.5- 1.0
- Page 119 and 120: 5 4 3 2 1 0! 8- 7- 6- 5- 4- 3- 0) E
- Page 121 and 122: Chapter , -11111 f'ol-ImItio I alld
- Page 123 and 124: (711 aI ý11 ýý r -', and dc\ -ý
- Page 125 and 126: Stability )I matioll wid dn cloplll
- Page 127 and 128: ( hapler, " C 101? 11 11 Confirrain
- Page 129 and 130: ( llý! pwl 3,1 I nuti 1Th III Unde
- Page 131 and 132: Table 3.3. Summary of time for chan
- Page 133 and 134: 14a 1) o rato 1 IM ("Al A1011 iIII(
- Page 135 and 136: Chapjý, i 4_ itito Trliclopllvwf,
- Page 137 and 138: Ch Therefore, an advantage of fluor
- Page 139 and 140: QWLu 1 nhorg() ,, týjjtioil into r
- Page 141 and 142: F-- I U V u 1 1 ý4 Q) ý4 Q. ) ý4
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Cha ei 4 I, ahoratorv i tj I. ItIO
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iloo m ci( pllvtohý: Ilthiý Illi
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4.4.5. Aims Laboralol v III cSti-ga
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Chqpiut 4 Lahorator\ ]m C. ý, tiua
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4.2.5. Light attenuation co-efficie
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Chaptýýi 4 [.. aboratorý )mcst .
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Chapici 4 1. ý)borator. \ imctigmi
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0.8 0.6 E 0.4 0.2 U) p 0 Z) 0.0 LL
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41 ahoralor\ imestipation itito n i
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CN cm ci 35n 30 25 20 15 10 5 0" 0.
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6-11 Lill Y-- CD 0 E I E 2i 300 250
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--I Iho-ratol\ illv, stilýatioll C
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E E m T 160- 140- 120- 100 cl, 80 -
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_0 0- x CD R) a) _c: 0) (D -C 2F) \
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Chapi(-r 4 ''1;, ýj()pl]ll tohcr)0
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C5 cq (q II: N C) Cl! I* (q 000o09C
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1 L) C: a) L) 0 CD CY) E =1 E x co
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(liaptei 4 k1borator, im cst4aatiol
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I. aborator\ jnvcstjýjtjojj jillo
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I lipt 41 ihra ' tjdi um flu iu Ion
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liapio--4 '(ýtl IýItloll into ill
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il-,, conditions, therefore elimina
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4.5 Acknowledgments 1 aborator" ilr
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In sit Iii TI t 1( )n Into -opilyl
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5.1.2. Aims The aims of the work we
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5.2.2. Colne Estuary (U. K. ) WU i'
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sim illvcstwaliorl into 1111crophyt
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)Icr -_-q ------ - ------ /H Sillý
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'E Low tide exposure period 1400- -
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0 E 0 E LL LL LL 0- C) c (5 5 -3 70
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0 a- -a D c E ýb CL 2000,1 (A) 150
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2000 - 1500- E looo- LL il 500- 01-
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,0 a- -a D c E CL 2000 - 1500- 1000
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_ln sim i lvesliganon into microphy
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24 22 20 18 a- E 14 E 12 10 2000- 1
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In Sim illv, -ýsliý, atloll imo l
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Liý- E (D L) 0 0.75- 0.70- 0.65- 0
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Table 5.6: Summary of Fo 15 data fo
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LO 1200 1100 1000 goo- 800- T 700-
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In sjtu-1 I'l ýcý, j I. 2allopint
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increasing PPFD and temperatures as
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will lllvcý, IIL 73 gMol M-2 s-1.
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In wflý ijlvt:,,, tw'allor) wto nl
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Im'C"OLattorl lilto lnljýl' Holl l
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ý! Yc Jill Lýla[mn 5) Cells from
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vallutý-orvý injil; - Microscale
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Ila Fllý 11 hiofihný, dominated s
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6.1.2. Investigating the sediment l
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Table 6.1: Light attenuation co-eff
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llýA )1ýýl 0 6.1.3. Examining pr
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- 6.1.4. Aims of investigation The
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( Ht: () ('ic ii ! t1)' (B) Samplin
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Chapm 6 Dicl \anjtions in rnicrophv
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C. ) 4-I 0 C. ) i H 9b Ln cd od CIS
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0 D 8.1 CS m CD CD 000C, 0 12 0 0 0
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I-j at io ri's iIT III C ]'Ot ýl ]
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C) N C> 0 CD CD 0 C, Cl) C-4 CD CD
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t2- C) C) CD C) 0 C) CD (0 u2 ýt (
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F CY) CF) co (D cu (D C) _0 12 U) c
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;3 C) 0 CN cu R (D 7, CD cu ý4 (D
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( hapLi i)cla ILt)H in. rnieiopIi E
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Figure 6.9: Change in taxonomic com
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volt W%ýq 001 Figure 6.11: It is e
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Light attenuation co-efficients (Ju
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g Mi MS MS I r. IIII-iI r--r--Tý I
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( l1q)lLi () Dfti 'L at ri nn' a ma
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III II "c I iiiii 7-1 IN, C, PPFD I
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hqplel 0 Table 6.9: Summary of mean
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cö cm E 0 cm E 12- E .r 3 68 10 12
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100 80 60 a E 40 -0 151 20 Arlesfor
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( h)t! 6 Dicl arljnoilý ill nlicro
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( 1ki l)ict ji roph bcnthi - hofli1
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( 1.1 )ic! anJtR)r1 fl flhICIuph\ i
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Chapter 6 Diel variations in microp
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Dicl varijtion, ý in microphytobcw
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Chapici_6 ___ -- - '- ---, -- -f),
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Did val ljlilor), ýIill rI, II 3)
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Mwiatorv -- ------ - ------- -;. no
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iý-, ratol S. 11011-1311, ýI-JIOF
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1-yýiwiv vs. non-mi that cells can
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Chapel - but also because cells can
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( ILtplet \1 alui ion mip2i1t)r in
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( Iiq»i 4. nor miiI 7.2.2. Experim
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Io fil I li I< samples) and maximum
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ýý ""j"l-4 placed in the I O'C wa
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. 1t. r- ,E - - 4 0 . - =UU cn CZ 9
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weaq 5uqejr4es E)L# 5ulinp plalA eo
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Ch MiLratoiv Hypothesis 3: Exposure
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1) 0 0 rdD 0 0 0 Cý3 4. u VI vI "c
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-0 700- 650- 600- 550- L) 500. C: u
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Chapicr 7 Nli-ratovv ý,,, non- nig
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CllaL)t i, -, ratory vs. no V biol-
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far-red conditions, demonstrating a
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Migratory vs. affect the fluorescen
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Mioiatovv \",,. mm-inioi-atory hiot
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( IIph. F \f atul . flI1 al()rv hio
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'\41_ý'Iatorv vs. non-illiorator"
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Chý N"I blofilllllýý artificial
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Mj-,, ralolv Vs. 11011-Irl -a orv b
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- -- --i;.. H)111fl bih1ni adaptati
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\1igraior '. oi-nii tor hhiiii' sed
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7.5. Acknowledgments '12I ttor \ '.
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8. General Discussion (Icncral Disc
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I--- 11 -- I- -- ( icricrid I estua
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(- II ap týý Iý8- -- ---- (icncr
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Ctlai,! ': ý ý, -II( Icticral Dis
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Chaoter S General Discussion the ef
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(IM The balance between all these p
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8.3.2. Cryolanding Overestimated be
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8 -( icticrulDiscu,, study). The ch
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Tide driven Black box C02 limitatio
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REFERENCES c Icrci wes
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Blanchard, G. F., Guarini, J-M., Ri
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Delo, E. A. and Ockenden, M. C. (19
- Page 372 and 373:
I AO'Clo! -, , Harper, M. A. (1977)
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photosynthetic communities: Fibre-o
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environment. Biopress, Bristol. (38
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G., McGrorty, S. and Eastwood, J. A
- Page 380 and 381:
Round, F. E. and Eaton, J. W. (1966
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1, TI.! -1, Paterson, D., M. (1999)
- Page 384:
Yentsch, C. S. and Yentsch, C. M. (