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Pharmaceutical botany - Lighthouse Survival Blog

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FERTILIZATION 49<br />

liantly colored corollas, have fragrant odors, and secrete nectar, a sweet<br />

liquid very attractive to insects which are adapted to this work through<br />

the possession of a pollen-carrying apparatus. Ex. : Orchids.<br />

Plants pollinated through the agency of water currents<br />

are known as Hydrophilous. To this class belong such plants as<br />

live under water and which produce flowers at or near the surface of<br />

the same. Ex. : Sparganium.<br />

Some plants as the Honeysuckle and Nasturtium are fertilized by<br />

humming birds.<br />

Before the pollen grain has been deposited upon the stigma a series<br />

of events affecting both the pollen grain and the embryo sac occur.<br />

The microspore (pollen grain) divides into two cells, the mother and<br />

tube cells of the male gametophyte. The nucleus of the mother<br />

cell divides to form two generative nuclei. The nucleus of the megaspore<br />

or embryo sac undergoes division until eight daughter nuclei<br />

are produced which are separated into the following groups<br />

(a) Three of these nuclei occupy a position at the apex, the lower<br />

nucleus of the group being the egg or ovum, the other two nuclei being<br />

the SYNERGiDS or assisting nuclei.<br />

(b) At the opposite end of the sac are three nuclei known as the<br />

antipodals which apparently have no special function.<br />

(c) The two remaining nuclei (polar nuclei) form a group lying<br />

near the centre of the embryo sac which unite to form a single nucleus<br />

from which, after fertilization, the endosperm of nourishing material<br />

is derived. This stage of the embryo sac constitutes the female<br />

gametophyte.<br />

Fertilization.—After the pollen grain reaches the stigma the viscid<br />

moisture of the stigma excites the outgrowth of the male gametophyte<br />

which bursts through the coats of the pollen grain forming a pollen<br />

tube. The pollen tube carrying within its walls two generative and<br />

one tube nucleus penetrates through the loose cells of the style until it<br />

reaches the micropyle of the ovule, then piercing the nucellus it enters<br />

the embryo sac. The tip of the tube breaks and one of the generative<br />

nuclei unites with the egg to form the oospore. The oospore develops<br />

at once into an embryo or plantlet, which lies passive until the seed<br />

undergoes germination. The other generative nucleus unites with the<br />

previously fused polar nuclei to form the endosperm nucleus which soon<br />

undergoes rapid division into a large number of nuclei scattered about<br />

through the protoplasm of the embryo sac. These accumulate proto-

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