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ECHINODERMATA - KU ScholarWorks - University of Kansas

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BLASTOID STUDIES 75<br />

Genus HETEROBLASTUS Etheridge & Carpenter,<br />

1886<br />

Type-species, by original designation.—Heteroblastus cumberlandi<br />

ETHERIDGE & CARPENTER, 1886.<br />

Generic diagnosis.—Spiraculate blastoids with nine<br />

spiracles, or eight spiracles and anispiracle, apparently<br />

with anispiracle located between epideltoid and hypodeltoid,<br />

lancet covered by side plates, probably more<br />

than one fold on each side <strong>of</strong> an ambulacrum, with<br />

one pore between side plates, radials overlapped by<br />

deltoids, deltoids long, with high coronal processes,<br />

and calyx form elliptical in side view. Lower Carboniferous,<br />

England.<br />

Remarks.—The genus Heteroblastus may have<br />

been derived from a form similar to Cri broblastus, in<br />

which the deltoids migrated aborally and overlapped<br />

the radial limbs.<br />

144<br />

D<br />

HETEROBLASTUS CUMBERLANDI<br />

Etheridge & Carpenter<br />

Plate 39, figures 1-2; text-figs. 144-146<br />

Heteroblastus cumberlandi ETHERIDGE & CARPENTER, 1886, pp. 141,<br />

237, ol. 6, figs. 1-6.<br />

Description.—Calyx calcitic, fragmentary, crushed, 10<br />

mm. long by 7 mm. wide, with base destroyed. Radials<br />

each 4 mm. long by 3 mm. wide, deltoids each 7 mm.<br />

long by 2.5 mm. wide, overlapping radials. Ambulacra<br />

five, long, linear, each 9 mm. long by 1.5 mm. wide, with<br />

lancet covered by side plates, 30 side plates in space <strong>of</strong><br />

10 mm. along an ambulacrum, and one pore between<br />

plates. Two small spiracles are excavated in the adorai<br />

tips <strong>of</strong> each deltoid, one spiracle on either side <strong>of</strong> the base<br />

<strong>of</strong> the high coronal process. Side plates appear to be<br />

normally disposed, with an outer side plate on the abmedial-adoral<br />

corner <strong>of</strong> each primary side plate, and<br />

pores present along deltoid and ambulacral margins. The<br />

remainder <strong>of</strong> the generic diagnosis was taken from<br />

ETHERIDGE & CARPENTER (1886).<br />

Occurrence.—Lower Carboniferous, Yoredale Shale,<br />

near Hexham, Northumberland, England.<br />

Type.—Topotype, S3,776, one specimen, Springer collection,<br />

U. S. National Museum, Washington.<br />

Genus LOPHOBLASTUS Rowley, 1901<br />

Type-species, by original designation.—Codonites inopinatus Row-<br />

LEY & HARE, 1891.<br />

Generic diagnosis.—Spiraculate blastoids with nine<br />

spiracles, or eight spiracles and an anispiracle which<br />

is located between an ?epideltoid and hooded hypodeltoid,<br />

two cryptodeltoids may be present, three<br />

hydrospire folds on each side <strong>of</strong> an ambulacrum, lancet<br />

exposed one-third <strong>of</strong> its width, with one pore between<br />

adjacent side plates along deltoid and radial<br />

margins, deltoids overlapping radials; shape ellipsoidal.<br />

Mississippian, Missouri.<br />

FIGURES 144-146. Heteroblastus cumberlandi ETHERIDGE<br />

& CARPENTER, Lower Carboniferous, England (U.S. Natl.<br />

Mus. no. S3,776). 144. Adoral tip <strong>of</strong> deltoid, side<br />

view showing elongate pores, X37.5. 145. Cross section<br />

<strong>of</strong> ambulacrum, X37.5. 146. Part <strong>of</strong> ambulacrum,<br />

showing side plates, X37.5. [Explanation.—Bp,<br />

brachiolar pit; C, canal (radial); D, deltoid; L, lancet;<br />

OSp, outer side plate; P, pore; S, spiracle; Sp, main side<br />

plate.]<br />

Remarks.—The genus Lophoblastus may have been<br />

derived from a form similar to Schizotremites that<br />

became elliptical in shape and developed nine definite<br />

spiracular openings, with reduction <strong>of</strong> hydrospire<br />

folds to three.<br />

LOPHOBLASTUS INOPINATUS (Rowley & Hare)<br />

Plate 45, figures 1-9; text-figs. 147-150<br />

Codonites inopinatus ROWLEY & HARE, 1891, p. 100, pl. 2, figs. Il-<br />

12; p. 118, pl. 3, fig. 17.<br />

Description.—Calyx subellipsoidal, 8.5 mm. long by<br />

6.5 mm. wide, with rounded summit, convex base, periphery<br />

at mid-height, vault 7 mm. long, pelvis 1.5 mm. long,<br />

and pelvic angle 120 degrees. Stem round, small, crenellar,<br />

about 1 mm. in diameter, attached to a raised pentagonal<br />

area <strong>of</strong> the basal circlet, which is convex, pentagonal<br />

in basal view, 1 mm. long by 2.5 mm. wide, with azygous<br />

basal broadly quadrangular and other two basais pentagonal,<br />

each with fine growth striae parallel to plate mar-

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