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ECHINODERMATA - KU ScholarWorks - University of Kansas

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64 THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS<br />

by side plates at aboral end and gradually exposed to<br />

one-third <strong>of</strong> its width near the adorai end, with one<br />

pore between side plates along deltoid and radial margins;<br />

radials overlapping deltoids; calyx subglobular.<br />

Mississippian, Illinois, Missouri, Kentucky, Alabama.<br />

Remarks.—The genus Diploblastus was probably<br />

derived from a form similar to Schizotremites in<br />

which the lancet plates extended downward and the<br />

form became subglobular. There should be more intermediate<br />

forms.<br />

DIPLOBLASTUS GLABER (Meek & Worthen)<br />

Plate 48, figures 1-12; plate 49, figures 1-9; text-figs. 113-119<br />

Granatocnnus glaber MEEK & WORTHEN, 1869, p. 91.<br />

Granatocrinus glaber MEEK & WORTHEN, 1873, p. 537, pl. 20, fig.<br />

I l.<br />

Description.—Observations here recorded are based<br />

mainly on one <strong>of</strong> the original syntypes. Calyx calcitic,<br />

worn, flattened spheroidal in side view, rounded pentagonal<br />

in top view, 5 mm. long by 6 mm. wide, with<br />

periphery at mid-height. The stem is round, crenellar,<br />

attached to a raised hexagonal base in the middle <strong>of</strong> the<br />

basal circlet, with small round central lumen. Basal circlet<br />

pentagonal in basal view, flat, 2.75 mm. in diameter, with<br />

three normally disposed basals. In one specimen from<br />

Kentucky the azygous basal is in the "DE" instead <strong>of</strong><br />

"AB" position, but this specimen may belong to another<br />

species. Radials five, pentagonal in side view, recurved<br />

below, each 4.5 mm. long by 3.25 mm. wide, with long,<br />

narrow, shallow sinus 4.5 mm. long by 1 mm. wide.<br />

Radials overlap deltoids.<br />

Deltoids four, short, each 2 mm. long by 2 mm. wide,<br />

broadly lanceolate, each with a paired spiracle at the<br />

adorai tip. Four deltoid plates are present on the anal<br />

side—a superdeltoid, two cryptodeltoids, and a hypodeltoid.<br />

The V-shaped superdeltoid is adjacent to the<br />

oral opening and has two deltoid septa that are continuous<br />

in position with the cryptodeltoids. The adorai<br />

end <strong>of</strong> the paired anispiracle is excavated in the superdeltoid.<br />

Immediately aboral to the two septa and aligned<br />

with them are two subquadrangular cryptodeltoids, located<br />

on either side <strong>of</strong> the anal opening. The broad<br />

pentagonal hypodeltoid rests on the cryptodeltoids and<br />

covers the aboral part <strong>of</strong> the anal opening. The anal<br />

opening is partly fused with the adjacent hydrospire<br />

canals, thus forming a paired anispiracle on the anal<br />

side. Therefore, five spiracles surround the oral opening<br />

—four paired spiracles and a paired anispiracle. In<br />

weathered specimens these openings are separate and a<br />

variable number appear to be present from specimen to<br />

specimen. There are two hydrospire folds on each side <strong>of</strong><br />

an ambulacrum, without a hydrospire plate.<br />

Ambulacra five, linear, recurved below, each 7.5 min.<br />

long by 0.75 mm. wide, with the median one-third <strong>of</strong><br />

the lancet exposed near the adoral end, but near the<br />

aboral end the lancet is covered by the side plates. There<br />

are 45 side plates in a space <strong>of</strong> 10 mm. along an ambulacrum.<br />

Th primary side plates are dumbbell-shaped with<br />

one large pore between adjacent plates at the abmedial<br />

margin <strong>of</strong> the lens-shaped outer side plate, with pores<br />

located along radial and deltoid ambulacral margins. The<br />

surfaces <strong>of</strong> the basal and radial plates are ornamented<br />

with fine growth lines parallel to plate margins and the<br />

deltoids have aligned granules along growth ridges.<br />

Remarks.—There is regional and stratigraphic variation<br />

among various specimens assigned to this species.<br />

The basal circlet is convex, flat, or slightly concave in<br />

different specimens, and on this basis alone one may<br />

distinguish various species. The object <strong>of</strong> the present<br />

work is to define genera, and therefore this problem will<br />

have to be considered by later investigators. The variation<br />

in shape <strong>of</strong> specimens from the St. Louis Group <strong>of</strong> Illinois<br />

and the Ste. Genevieve Limestone <strong>of</strong> Kentucky<br />

should be called to attention.<br />

Occurrence.—Mississippian, St. Louis Limestone, Hardin<br />

County, Illinois (syntypes, X-363) and Brown<br />

County, Illinois (E1,498). Ste. Genevieve Limestone,<br />

Crittenden County, Kentucky (1,303). Gasper Formation,<br />

Huntsville, Alabama (71,650). Warsaw Formation,<br />

Florence, Alabama (S3,782).<br />

Types.—Syntypes, X-363, three specimens, one 4-<br />

sided, A. H. Worthen collection, old Illinois State Museum<br />

no. 10,096, now in Department <strong>of</strong> Geology, <strong>University</strong><br />

<strong>of</strong> Illinois, Urbana. Plesiotype, E1,498, one specimen,<br />

labelled Mesoblastus kirkwoodensis, A. H. Worthen<br />

collection, <strong>University</strong> <strong>of</strong> Illinois. Plesiotypes, 1,303, seven<br />

specimens, one polished section, one thin section, and five<br />

EXPLANATION OF PLATE 17<br />

FIGURE PAGE<br />

1-3,7-9. Astrocrinus benniei (ETnERIDGE), plesiotypes, 362,<br />

Harvard Mus. Comp. Zoology, Lower Carboniferous Limestone<br />

Group (shale above no. 2 limestone), Carlops Quarry,<br />

Peebleshire, Scotland; 1,3, oral and aboral views with<br />

"A" ambulacrum toward top <strong>of</strong> page in fig. / ( X 10.6) ;<br />

2, 9, "C" ambulacral and aboral views <strong>of</strong> another specimen<br />

(approximately X14), 7,8, aboral and oral views <strong>of</strong><br />

specimen with "D" side directed downward ( X17.0) 21<br />

4-6. Pleuroschisma ontario REIMANN, topotype, 35,033, Southworth<br />

coll., Univ. Michigan; Devonian, Hungry Hollow<br />

Formation (coral bed), Ausable River valley northeast <strong>of</strong><br />

Arkona, Ontario; oral, "D" ambulacral, aboral views (all<br />

X9.1) 42

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