ECHINODERMATA - KU ScholarWorks - University of Kansas
ECHINODERMATA - KU ScholarWorks - University of Kansas
ECHINODERMATA - KU ScholarWorks - University of Kansas
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62 THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS<br />
DEVONOBLASTUS LEDA (Hall)<br />
Plate 24, figures 1-2, 6<br />
Pentrenntes leda HALL, 1862, p. 149, pl. 1, fig. 11.<br />
Devonoblastus leda REIMANN, 1935, p. 31.<br />
Description.—A detailed description <strong>of</strong> the syntypes<br />
has been submitted for publication in another work. The<br />
syntypes are not well preserved and most <strong>of</strong> the generic<br />
information is taken from the description <strong>of</strong> specimens<br />
<strong>of</strong> Devonoblastus whiteavesi. The syntypes are figured on<br />
Plate 24. This species differs from D. whiteavesi in that<br />
the maximum width (or periphery) <strong>of</strong> D. leda is low,<br />
at or near the radial lips, in contrast to its higher, more<br />
median position in D. whiteavesi.<br />
Occurrence.—Devonian, Hamilton Shale, western<br />
New York.<br />
Types.—Syntypes, 451 (figures 1, 6), and 452 (figure<br />
2), two specimens, Hall collection, New York State Museum,<br />
Albany, New York.<br />
DEVONOBLASTUS WHITEAVESI Reimann<br />
Plate 24, figures 3-5; plate 25, figures 1-12; text-figs. 101-112<br />
Devonoblastus whiteavesi REIMANN, 1935, p. 32, pl. 1, fig. 8.<br />
Granatocrinus leda WHITEAVES, 1889, p. 108, pl. 14, fig. 14.<br />
Description.—The description is mainly on specimen<br />
35,036, supplemented by information from other specimens.<br />
The calyx is calcitic, oval in side view, with short<br />
conical pelvis, 16 mm. long by 10.5 mm. wide, pentagonal<br />
in top view, with periphery near mid-height, and vault<br />
14 mm. long, pelvis 2 mm. long, pelvic angle on basais<br />
85 degrees and on radial bodies 130 degrees. The stem<br />
is round, crenellar, 1.5 mm. in diameter, with approximately<br />
45 crenellae extending inward radially from the<br />
margin one-third <strong>of</strong> the distance to the small round<br />
lumen. The stem is attached to the basal circlet, which<br />
has a hexagonal raised area at the aboral ends <strong>of</strong> the basal<br />
plates. Basal circlet pentagonal in aboral view, widely<br />
conical in side view, 2 mm. long by 5 mm. wide, with<br />
strongly rounded medial ridge on each basal. Radials five,<br />
elongate hexagonal, each 14.5 mm. long by 5.5 mm.<br />
wide, with long, narrow, shallow sinus 13 mm. long by<br />
2 mm. wide; radials overlapping deltoids. The surfaces<br />
<strong>of</strong> the radials and basais are ornamented by fine growth<br />
striae parallel to plate margins.<br />
Deltoids four, lancet-shaped, each 3.5 mm. long by<br />
2 mm. wide, visible in side view, with one large oval<br />
spiracle notched in the adorai end <strong>of</strong> each, separating the<br />
deltoid into two parts, the deltoid lip located adjacent to<br />
the oral opening and the deltoid body adjacent to the<br />
radial limbs. The surfaces <strong>of</strong> the deltoid bodies are ornamented<br />
by coarse growth ridges parallel to plate margins,<br />
especially parallel to the radiodeltoid sutures. On the anal<br />
side four anal deltoid plates seem to be present. The small<br />
pentagonal superdeltoid is adjacent to the oral opening<br />
and forms the external border on the adorai part <strong>of</strong> the<br />
large oval anispiracle. Internally the two cryptodeltoids<br />
rest on the aboral face <strong>of</strong> the superdeltoid and separate<br />
the anal opening from the adjacent hydrospire canals.<br />
The cryptodeltoids are infolded to form hydrospires on<br />
the anal side, and are overlapped by the radial limbs. The<br />
cryptodeltoids are covered externally by the hypodeltoid<br />
plate and are not visible from the exterior. The pentagonal<br />
hypodeltoid is visible in side view and is homologous<br />
with the deltoid body <strong>of</strong> each <strong>of</strong> the other four deltoids.<br />
Five hydrospire folds are seen on each side <strong>of</strong> an<br />
ambulacrum.<br />
Ambulacra five, linear, slightly recurved below periphery,<br />
each 16 mm. long by 2 mm. wide, with lancet covered<br />
by side plates except at the adorai end, 24 side plates<br />
in a space <strong>of</strong> 10 mm. along an ambulacrum. The side<br />
plates are normally disposed, with four main cover-plate<br />
lobes to each side plate along the main food groove. A<br />
single large pore is observed between adjacent side plates<br />
along the radial ambulacral margins, and one or two<br />
pores occur along the deltoid ambulacral margins just<br />
above the radiodeltoid sutures. Each spiracle is bordered<br />
by a deltoid lip, two lancet stipes, two side plates, and a<br />
deltoid body.<br />
Occurrence.—Middle Devonian, coral zone <strong>of</strong> Widder<br />
beds, Hungry Hollow Formation, Thedford and Arkona<br />
areas, Ontario. Shale layer <strong>of</strong> Tichenor Limestone, Cazenovia<br />
Creek, Springbrook, Erie County, New York.<br />
Types.—Topotype, 35,036, one specimen, Charles<br />
Southworth collection, from Ausable River valley, 2.5<br />
miles northeast <strong>of</strong> Arkona; <strong>University</strong> <strong>of</strong> Michigan, Ann<br />
Arbor. Plesiotype, 34,467, one specimen, from tile yard<br />
0.5 mile north <strong>of</strong> Thedford, Ontario, Charles Southworth<br />
collection (June, 1954), <strong>University</strong> <strong>of</strong> Michigan. Plesiotypes,<br />
E13,065, three specimens, Charles Southworth collection<br />
(1941), from area near Thedford, Ontario, Buffalo<br />
Society <strong>of</strong> Natural Sciences, Buffalo, New York. Hypotype,<br />
3,661, one specimen, from Thedford area, described<br />
by WHITEAVES (1889) as Granatocrinus leda, Geological<br />
Survey <strong>of</strong> Canada, Ottawa, Ontario, Canada. Plesiotypes,<br />
S4,648, two specimens, Springer collection, from Ausable<br />
River valley, 2 miles east <strong>of</strong> Arkona, Ontario, collected<br />
by Charles Southworth; S4,144, one specimen, Springer<br />
collection, from Marsh's Mill, Arkona, Ontario, lower<br />
Widder beds; and 142,025, one specimen, labelled Devonoblastus<br />
leda, Tichenor Limestone, New York, all in<br />
U. S. National Museum, Washington.<br />
Genus DEPLOBLASTUS Fay, n. gen.<br />
Type-spedes, by original designation (herein).—Granatocrinus glaber<br />
MEEK & WORTHEN, 1869.<br />
Generic diagnosis.—Spiraculate blastoids with five<br />
paired spiracles, or four paired spiracles in addition to<br />
a paired anispiracle, with superdeltoid, two cryptodeltoids,<br />
and hypodeltoid on anal side; two hydrospire<br />
folds on each side <strong>of</strong> an ambulacrum; lancet covered