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ECHINODERMATA - KU ScholarWorks - University of Kansas

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BLASTOID STUDIES 89<br />

with center at junction <strong>of</strong> interradial suture with the<br />

radiodeltoid suture.<br />

Ambulacra five, linear, with lancet exposed along the<br />

median one-third <strong>of</strong> its width almost to the aboral end<br />

<strong>of</strong> each ambulacrum. Side plates normally disposed, with<br />

approximately five cover-plate sockets to each side plate<br />

along main food groove, and two pores to each side<br />

plate, one <strong>of</strong> which is almost even with a side plate<br />

suture and the other about midway between the side<br />

plate sutures. Surfaces <strong>of</strong> calyx plates ornamented by<br />

fine granules arranged subparallel to plate margins along<br />

growth lines.<br />

Occurrence.—Lower Carboniferous, recorded as Devonshire<br />

(may mean Derbyshire), England.<br />

Type.—Plesiotype, 1,633, one specimen, James collection,<br />

labelled Granatocrinus ellipticus, Walker Museum,<br />

<strong>University</strong> <strong>of</strong> Chicago.<br />

ORBITREMITES ELLIPTICUS (Sowerby)<br />

Plate 43, figures 1-3, 10-11; text-figs. 186, 187<br />

Pentremites elliptica SOWERBY, 1825, p. 317, pl. 11, fig. 4.<br />

Elaeacrinus ellipticus SHUMARD, 1863, p. 112.<br />

Description.—Calyx calcitic, fragmentary, 12.5 mm.<br />

long by 12 mm. wide, ellipsoidal in side view, rounded<br />

pentagonal in top view, with flat summit, and periphery<br />

above mid-height. Stem and basais destroyed. Radials<br />

five, elongate hexagonal in side view, reaching adorally<br />

to periphery with narrow, shallow sinus 10 mm. long by<br />

1.5 mm. wide. Deltoids overlap radials. The angle <strong>of</strong><br />

the radial limbs at the radiodeltoid suture, with center<br />

at interradial suture is 85 degrees.<br />

Deltoids four, large, broadly rhombic, reaching below<br />

periphery, each 8 mm. long by 5.5 mm. wide, with one<br />

large triangular spiracle in the middle <strong>of</strong> the depressed<br />

adorai tip. On the anal side the triangular anispiracle is<br />

located between a pentagonal superdeltoid and a pentagonal<br />

hypodeltoid, with two hidden cryptodeltoids. Thus,<br />

five openings ocur around the oral aperture, consisting <strong>of</strong><br />

four spiracles and an anispiracle. There is one hydrospire<br />

fold on each side <strong>of</strong> an ambulacrum, terminating<br />

admedially in a hydrospire plate.<br />

Ambulacra five, linear, raised, each 16 mm. long by<br />

1.5 mm. wide, with lancet exposed along the middle onethird<br />

<strong>of</strong> its width almost to the aboral end, and 26 side<br />

plates in a space <strong>of</strong> 10 mm. along an ambulacrum. There<br />

are two pores to each side plate along the deltoid and<br />

radial margins. The surfaces <strong>of</strong> the calyx plates are almost<br />

smooth, with fine growth striae subparallel to plate<br />

margins, and granular raised ridges immediately aboral<br />

to each spiracle and anispiracle.<br />

Occurrence.—Lower Carboniferous, upper part <strong>of</strong><br />

Clitheroe Limestone (C 2) <strong>of</strong> Salt Hill Knoll, Clitheroe,<br />

Lancashire, England (Beaver collection), and Carboniferous<br />

limestone, Yorkshire, England (deKoninck collection).<br />

Types.—Plesiotypes, 245, 11 specimens, deKoninck<br />

collection, old numbers, 2,324 and 467, Harvard Museum<br />

<strong>of</strong> Comparative Zoology, Cambridge, Mass.; one unnumbered<br />

specimen, Beaver collection, sent by DR. JOYSEY from<br />

Cambridge, England, Humble Oil & Refining Co., Houston,<br />

Texas. The above description is based on the specimen<br />

in the Beaver collection.<br />

Genus PENTREMITES Say, 1820<br />

Type-species, by subsequent designation (ETHERIDGE & CARPENTER,<br />

1886).—encrina Godonii DEFRANCE, 1819.<br />

Generic diagnosis.—Spiraculate blastoids with five<br />

spiracles or four spiracles and an anispiracle, with the<br />

anispiracle excavated in one anal deltoid plate, ?two,<br />

three to seven or more hydrospire folds on each side<br />

<strong>of</strong> an ambulacrum, oral and anal areas covered by<br />

many imbricate plates, one pore between side plates<br />

along deltoid and radial margins, lancet almost completely<br />

exposed, ambulacra petaloid, radials overlap<br />

deltoids, and shape <strong>of</strong> calyx in side view rounded obconical<br />

or club-shaped. Mississippian -Pennsylvanian,<br />

Alabama, Tennessee, Kentucky, Indiana, Illinois, Missouri,<br />

Iowa, Oklahoma, Arkansas, Mississippi, Montana,<br />

Alberta, ?Alaska.<br />

Remarks.—The genus Pentremites was probably<br />

derived from Devon oblastus by fusion <strong>of</strong> the anal<br />

deltoid plates into one, and continued downward<br />

migration <strong>of</strong> the deltoid plates, and outward migration<br />

<strong>of</strong> the lancet plates. A detailed description <strong>of</strong> the<br />

neotype (new holotype) <strong>of</strong> P. godoni is reserved for<br />

another paper to appear in the Journal <strong>of</strong> Paleontology.<br />

EXPLANATION<br />

FIGURE<br />

PAGE<br />

1-6. Elaeacrinus venustus (MILLER & GURLEY). 1 3. Syntype,<br />

6,114, Univ. Chicago; Devonian, Columbus, Ohio;<br />

oral, "D" ambulacral, aboral views <strong>of</strong> figured type (Miller<br />

& Gurley, 1894, pl. 8, fig. 26-30) (all X4.1).-4-6.<br />

Syntype, 1,317, Univ. Cincinnati; Devonian, Columbus,<br />

OF PLATE 28<br />

Ohio; oral, "D" ambulacral, aboral views (all X3.7) 69<br />

7-10. Elaeacrinus verneudi ROEMER, plesiotype, 9,956, Gurley<br />

coll., Univ. Chicago; Devonian, Jeffersonville Limestone,<br />

Jeffersonville, Ind., 7-9, oral, aboral, "D" ambulacral views<br />

( X1.7); 10, enlarged oral view <strong>of</strong> summit ( X4.3) 68

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