ECHINODERMATA - KU ScholarWorks - University of Kansas
ECHINODERMATA - KU ScholarWorks - University of Kansas
ECHINODERMATA - KU ScholarWorks - University of Kansas
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84 THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS<br />
side <strong>of</strong> an ambulacrum with short hydrospire canal;<br />
form <strong>of</strong> calyx widely elliptical. Devonian, New York,<br />
Iowa, Ontario, Michigan.<br />
Remarks.-The genus Nucleocrinus was probably<br />
derived from Elaeacrinus by supression <strong>of</strong> the superdeltoid<br />
plate internally and reduction in the number <strong>of</strong><br />
oral plates to five.<br />
NUCLEOCRINUS ELEGANS Conrad<br />
Plate 29, figures 4-6, 9; text-fig. 168<br />
Nucleocrinus elegans CONRAD, 1842, p. 280, pl. 15, fig. 17.<br />
Description.-Calyx calcitic, subglobular in side view,<br />
rounded pentagonal in top view, 8.5 mm. long by 11 mm.<br />
wide, with concave summit and concave base, and periphery<br />
above mid-height. Stem round, crenellar, 0.75 mm.<br />
in diameter, in basal concavity, with small central round<br />
lumen. Basal circlet pentagonal in basal view, 1.5 mm.<br />
wide, in basal concavity, with three normally arranged<br />
basals. Radials five, short, pentagonal in side view, each<br />
2.5 mm. long by 3.5 mm. wide, mostly on aboral surface,<br />
recurved below in basal concavity, with short, shallow<br />
sinus 1 mn-i, long by 1 mm. wide, and long flaring lips<br />
projected aborally into rounded ridges. Deltoids overlap<br />
radials. The surfaces <strong>of</strong> the basais and radials are ornamented<br />
by fine growth striae subparallel to plate margins.<br />
Deltoids four, flattened lenticular, each 8 mm. long<br />
by 6 mm. wide, each with a broadly curved V-shaped<br />
radiodeltoid suture at the base, and with two hemielliptical<br />
spiracles at the adorai tip along ambulacral margins.<br />
The ornamentation <strong>of</strong> the deltoids consists <strong>of</strong> two parts,<br />
a central triangular portion with apex at adorai end, and<br />
two bordering lenticular portions. The central part has<br />
fine growth ridges parallel to the radiodeltoid suture and<br />
the marginal portions have narrow nodes that are aligned<br />
with the adjacent side plates at a high angle to the<br />
ambulacra. On the anal side three deltoids are visible,<br />
one deltoid being hidden. The visible deltoids are the<br />
two lenticular cryptodeltoids marginal to the ambulacra,<br />
with a central hypodeltoid that is subquadrangular in<br />
shape. The hypodeltoid, on the aboral side <strong>of</strong> the diamond-shaped<br />
anal opening has growth striae parallel<br />
to the radiodeltoid suture and lateral margins, the central<br />
adorai portion being raised. The lenticular cryptodeltoids<br />
extend from the adorai and lateral sides <strong>of</strong> the anal opening<br />
to the radiodeltoid suture and have aligned nodes<br />
similar to those <strong>of</strong> the other four deltoids. The adorai tips<br />
<strong>of</strong> the cryptodeltoids abut against a large anal oral plate<br />
and each tip has a hemielliptical spiracle along its ambulacral<br />
margin. Thus, ten spiracles in addition to the anal<br />
opening occur around the oral aperture, which is covered<br />
by five large arrow-shaped oral plates, interambulacral<br />
in position. Internally, beneath the anal oral and between<br />
the adorai tips <strong>of</strong> the cryptodeltoids it is assumed that a<br />
supressed superdeltoid is present. Hydrospire folds presumably<br />
two on each side <strong>of</strong> an ambulacrum, with a<br />
short hydrospire canal.<br />
Ambulacra five, linear, recurved below, raised above<br />
calyx surface, each 12 mm. long by 1 mm. wide, with<br />
lancet covered by side plates, and 30 side plates in a space<br />
<strong>of</strong> 10 mm. along an ambulacrum. The side plates consist<br />
<strong>of</strong> a large raised subquadrangular primary side plate,<br />
with a large triangular secondary side plate on the adoralabmedial<br />
bevelled corner <strong>of</strong> the primary side plate, and<br />
an inner side plate on the admedial side <strong>of</strong> each pore,<br />
broadly lenticular in shape. Approximately four main<br />
cover-plate sockets to each side plate occur along the main<br />
food groove, and one pore is present between adjacent<br />
side plates along the radial and deltoid margins.<br />
Occurrence.-Middle Devonian, Hamilton beds, Moscow,<br />
New York.<br />
Type.-Topotype, 19,049, one <strong>of</strong> two specimens, one<br />
<strong>of</strong> which is Placoblastus lucina, Gurley collection, Walker<br />
Museum, <strong>University</strong> <strong>of</strong> Chicago.<br />
NUCLEOCRINUS MELONIFORMIS (Barris)<br />
Plate 29, figures 1-3, 7-8; plate 30, figures 1-9; plate 32, figure 9;<br />
text-figs. 169-178<br />
Elaeacrinus meloniformis BARRIS, 1883, p. 361; 1885-86, p. 91, pl.<br />
1, fig. 3.<br />
Nucleocrinus meloniformis THOMAS, 1924, p. 428, pl. 36, figs. 10-11.<br />
Description.-Calyx calcitic, 10 mm. long by 7.5 mm.<br />
wide, with periphery above mid-height, vault 10 mm.<br />
long, pelvis 2 mm. long in oblique basal view, with pelvic<br />
FIGURES 168-177. Species <strong>of</strong> Nucleocrinus CONRAD, Middle Devonian, New York, Ontario.<br />
168. N. elegans CONRAD, Hamilton Group, Moscow, N.Y.<br />
(Univ. Chicago, no. 19,049); part <strong>of</strong> "B" ambulacrum<br />
showing main, inner, and outer side plates, X45.<br />
169-177. N. meloniformis (BARRIS), Hungry Hollow Formation,<br />
Thedford and Arkona, Ont. (Geol. Survey Canada,<br />
no. 3,662a; Buffalo Soc. Nat. Sci., nos. E148, E15,939,<br />
E21,120); 169, summit and anal areas (no. 3,662a), X15;<br />
170, basal area (no. 3,662a), X15; 171, stem impression<br />
(no. 3,662a), X45; 172, cross section <strong>of</strong> "E" ambulacrum<br />
(no. E21,120), X15; 173, summit area drawn from polished<br />
section showing hidden superdeltoid (no. E148),<br />
X15; 174, part <strong>of</strong> weathered "E" ambulacrum, showing<br />
side plates (E15,939), X45; 175, part <strong>of</strong> "E" ambulacrum,<br />
showing side plates (no. E15,939), X45; 176, anal<br />
area, oral plates lacking (no. E21,120), X15; 177, part <strong>of</strong><br />
"B" ambulacrum, showing main, inner, and outer side<br />
plates (no. 3,662a), X75. [Explanation.-An, anal opening;<br />
Bf, brachiolar facet; Bp, brachiolar pit; "C," amb.;<br />
CR, cryptodeltoid; "D," amb.; D, deltoid; HD, hypodeltoid;<br />
ISp, inner side plate; L, lancet; 0, oral plate;<br />
OSp, outer side plate; P, pore; S, spiracle; Sp, main side<br />
plate; Su, superdeltoid; Z, azygous basal.]