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ECHINODERMATA - KU ScholarWorks - University of Kansas

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84 THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS<br />

side <strong>of</strong> an ambulacrum with short hydrospire canal;<br />

form <strong>of</strong> calyx widely elliptical. Devonian, New York,<br />

Iowa, Ontario, Michigan.<br />

Remarks.-The genus Nucleocrinus was probably<br />

derived from Elaeacrinus by supression <strong>of</strong> the superdeltoid<br />

plate internally and reduction in the number <strong>of</strong><br />

oral plates to five.<br />

NUCLEOCRINUS ELEGANS Conrad<br />

Plate 29, figures 4-6, 9; text-fig. 168<br />

Nucleocrinus elegans CONRAD, 1842, p. 280, pl. 15, fig. 17.<br />

Description.-Calyx calcitic, subglobular in side view,<br />

rounded pentagonal in top view, 8.5 mm. long by 11 mm.<br />

wide, with concave summit and concave base, and periphery<br />

above mid-height. Stem round, crenellar, 0.75 mm.<br />

in diameter, in basal concavity, with small central round<br />

lumen. Basal circlet pentagonal in basal view, 1.5 mm.<br />

wide, in basal concavity, with three normally arranged<br />

basals. Radials five, short, pentagonal in side view, each<br />

2.5 mm. long by 3.5 mm. wide, mostly on aboral surface,<br />

recurved below in basal concavity, with short, shallow<br />

sinus 1 mn-i, long by 1 mm. wide, and long flaring lips<br />

projected aborally into rounded ridges. Deltoids overlap<br />

radials. The surfaces <strong>of</strong> the basais and radials are ornamented<br />

by fine growth striae subparallel to plate margins.<br />

Deltoids four, flattened lenticular, each 8 mm. long<br />

by 6 mm. wide, each with a broadly curved V-shaped<br />

radiodeltoid suture at the base, and with two hemielliptical<br />

spiracles at the adorai tip along ambulacral margins.<br />

The ornamentation <strong>of</strong> the deltoids consists <strong>of</strong> two parts,<br />

a central triangular portion with apex at adorai end, and<br />

two bordering lenticular portions. The central part has<br />

fine growth ridges parallel to the radiodeltoid suture and<br />

the marginal portions have narrow nodes that are aligned<br />

with the adjacent side plates at a high angle to the<br />

ambulacra. On the anal side three deltoids are visible,<br />

one deltoid being hidden. The visible deltoids are the<br />

two lenticular cryptodeltoids marginal to the ambulacra,<br />

with a central hypodeltoid that is subquadrangular in<br />

shape. The hypodeltoid, on the aboral side <strong>of</strong> the diamond-shaped<br />

anal opening has growth striae parallel<br />

to the radiodeltoid suture and lateral margins, the central<br />

adorai portion being raised. The lenticular cryptodeltoids<br />

extend from the adorai and lateral sides <strong>of</strong> the anal opening<br />

to the radiodeltoid suture and have aligned nodes<br />

similar to those <strong>of</strong> the other four deltoids. The adorai tips<br />

<strong>of</strong> the cryptodeltoids abut against a large anal oral plate<br />

and each tip has a hemielliptical spiracle along its ambulacral<br />

margin. Thus, ten spiracles in addition to the anal<br />

opening occur around the oral aperture, which is covered<br />

by five large arrow-shaped oral plates, interambulacral<br />

in position. Internally, beneath the anal oral and between<br />

the adorai tips <strong>of</strong> the cryptodeltoids it is assumed that a<br />

supressed superdeltoid is present. Hydrospire folds presumably<br />

two on each side <strong>of</strong> an ambulacrum, with a<br />

short hydrospire canal.<br />

Ambulacra five, linear, recurved below, raised above<br />

calyx surface, each 12 mm. long by 1 mm. wide, with<br />

lancet covered by side plates, and 30 side plates in a space<br />

<strong>of</strong> 10 mm. along an ambulacrum. The side plates consist<br />

<strong>of</strong> a large raised subquadrangular primary side plate,<br />

with a large triangular secondary side plate on the adoralabmedial<br />

bevelled corner <strong>of</strong> the primary side plate, and<br />

an inner side plate on the admedial side <strong>of</strong> each pore,<br />

broadly lenticular in shape. Approximately four main<br />

cover-plate sockets to each side plate occur along the main<br />

food groove, and one pore is present between adjacent<br />

side plates along the radial and deltoid margins.<br />

Occurrence.-Middle Devonian, Hamilton beds, Moscow,<br />

New York.<br />

Type.-Topotype, 19,049, one <strong>of</strong> two specimens, one<br />

<strong>of</strong> which is Placoblastus lucina, Gurley collection, Walker<br />

Museum, <strong>University</strong> <strong>of</strong> Chicago.<br />

NUCLEOCRINUS MELONIFORMIS (Barris)<br />

Plate 29, figures 1-3, 7-8; plate 30, figures 1-9; plate 32, figure 9;<br />

text-figs. 169-178<br />

Elaeacrinus meloniformis BARRIS, 1883, p. 361; 1885-86, p. 91, pl.<br />

1, fig. 3.<br />

Nucleocrinus meloniformis THOMAS, 1924, p. 428, pl. 36, figs. 10-11.<br />

Description.-Calyx calcitic, 10 mm. long by 7.5 mm.<br />

wide, with periphery above mid-height, vault 10 mm.<br />

long, pelvis 2 mm. long in oblique basal view, with pelvic<br />

FIGURES 168-177. Species <strong>of</strong> Nucleocrinus CONRAD, Middle Devonian, New York, Ontario.<br />

168. N. elegans CONRAD, Hamilton Group, Moscow, N.Y.<br />

(Univ. Chicago, no. 19,049); part <strong>of</strong> "B" ambulacrum<br />

showing main, inner, and outer side plates, X45.<br />

169-177. N. meloniformis (BARRIS), Hungry Hollow Formation,<br />

Thedford and Arkona, Ont. (Geol. Survey Canada,<br />

no. 3,662a; Buffalo Soc. Nat. Sci., nos. E148, E15,939,<br />

E21,120); 169, summit and anal areas (no. 3,662a), X15;<br />

170, basal area (no. 3,662a), X15; 171, stem impression<br />

(no. 3,662a), X45; 172, cross section <strong>of</strong> "E" ambulacrum<br />

(no. E21,120), X15; 173, summit area drawn from polished<br />

section showing hidden superdeltoid (no. E148),<br />

X15; 174, part <strong>of</strong> weathered "E" ambulacrum, showing<br />

side plates (E15,939), X45; 175, part <strong>of</strong> "E" ambulacrum,<br />

showing side plates (no. E15,939), X45; 176, anal<br />

area, oral plates lacking (no. E21,120), X15; 177, part <strong>of</strong><br />

"B" ambulacrum, showing main, inner, and outer side<br />

plates (no. 3,662a), X75. [Explanation.-An, anal opening;<br />

Bf, brachiolar facet; Bp, brachiolar pit; "C," amb.;<br />

CR, cryptodeltoid; "D," amb.; D, deltoid; HD, hypodeltoid;<br />

ISp, inner side plate; L, lancet; 0, oral plate;<br />

OSp, outer side plate; P, pore; S, spiracle; Sp, main side<br />

plate; Su, superdeltoid; Z, azygous basal.]

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