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ECHINODERMATA - KU ScholarWorks - University of Kansas

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94 THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS<br />

five large arrow-shaped oral plates, each interambulacral<br />

in position, the superdeltoid and adjacent adoral tips <strong>of</strong><br />

the cryptodeltoids abutting against the large anal oral.<br />

The adorai tips <strong>of</strong> each deltoid are excavated by two semielliptical<br />

spiracles along ambulacral margins, and each<br />

adorai tip <strong>of</strong> each cryptodeltoid has one marginal slitlike<br />

spiracle, separate from the anal opening. Thus, 11 openings<br />

surround the oral aperture, consisting <strong>of</strong> ten spiracles<br />

and a separate anal opening. The calyx plates are ornamented<br />

as in Nucleocrinus. Two elongate hydrospires<br />

occur on each side <strong>of</strong> an ambulacrum, with an elongate<br />

hydrospire canal extending to each.<br />

Ambulacra five, linear, recurved below, each 55 mm.<br />

long by 2 mm. wide, with 23 side plates in a space <strong>of</strong> 10<br />

mm. along an ambulacrum, lancet covered by side plates,<br />

inner side plate large, and side plates disposed as in<br />

Nucleocrinus. There is an inward-projecting hornlike<br />

process on the inner medial wall <strong>of</strong> the hypodeltoid. Six<br />

cover-plate sockets to each side plate are observed along<br />

the main food groove.<br />

Occurrence.—Middle Devonian, Thunder Bay Limestone<br />

(or Partridge Point Formation), Partridge Point,<br />

near Alpena, Michigan.<br />

Types.— Topotypes?, E21,113, four specimens;<br />

E21,115, two specimens, Buffalo Society <strong>of</strong> Natural Sciences,<br />

Buffalo, New York. The description is taken from<br />

the characters mainly seen in specimens E21,115.<br />

PLACOBLASTUS ANGULARIS (Lyon)<br />

Plate 32, figure 8; plate 33, figures 1-9; plate 34, figures 1-9; textfigs.<br />

197-199<br />

Olivanites angularis LYON, 1857, p. 492, pl. 5, figs. 2, 2a, 2b.<br />

Nucleocrinus angularis LYON & CASSEDAY, 1859, p. 295.<br />

Nucleocrinus powelli REIMANN, 1935, p. 34, pl. 3, figs. 4-6.<br />

Description.—After careful comparison <strong>of</strong> the types<br />

<strong>of</strong> P. powelli with P. angularis I have been unable to find<br />

significant differences. The description is <strong>of</strong> a paratype<br />

(E9,058) <strong>of</strong> P. powelli.<br />

Calyx calcitic, 19 mm. long by 15.5 mm. wide, with<br />

flat summit, slightly concave base, periphery slightly adoral<br />

<strong>of</strong> mid-height, and interambulacral areas concave in<br />

oral view. Basal circlet pentagonal in aboral view, 3 mm.<br />

wide, flat to slightly concave, with round stem impression<br />

1.5 mm. wide, with small round central lumen, and<br />

three normally disposed basais. Radials five, each 5 mm.<br />

long by 5.5 mm. wide, recurved below in basal concavity,<br />

with sinus 1.5 mm. long by 1 mm. wide. The radiodeltoid<br />

suture is broadly M-shaped. Deltoids overlap radials.<br />

Deltoids four, bluntly lenticular, each 18 mm. long<br />

by 8.5 mm. wide, each with a smooth central triangular<br />

area with base almost full width <strong>of</strong> radiodeltoid suture,<br />

bordered laterally by two lenticular areas <strong>of</strong> nodose ornamentation,<br />

similar to those <strong>of</strong> Nucleocrinus. Each row <strong>of</strong><br />

nodes is aligned with a side plate, forming a high angle<br />

with the median food groove. The aboral tips <strong>of</strong> the<br />

nodose side regions extend aborally between each ambulacrum<br />

and radial limb, forming a short wedge. On the<br />

anal side there are at least three deltoids, the two elongate<br />

lenticular cryptodeltoids with the elongate subquadrangular<br />

hypodeltoid between, and probably a fourth (superdeltoid)<br />

between the adorai tips <strong>of</strong> the cryptodeltoids. The<br />

superdeltoid is covered almost completely by an anal oral<br />

plate in approximately the same position, but in some<br />

specimens there appear to be two plates fully exposed<br />

in this region. Thus, six or seven oral plates cover the<br />

summit, depending on interpretation <strong>of</strong> the plate that is<br />

adjacent to the adoral side <strong>of</strong> the anal opening. There are<br />

five large oral plates over the summit, each arrow-shaped<br />

and interambulacral in position, with one or two accessory<br />

oral plates on the anal side. At present, the interpretation<br />

here presented is that two plates occur between the<br />

FIGURE 197. Placoblastus angularis (LYON), Middle<br />

Devonian, Ludlowville Formation, Springbrook, New<br />

York (Buffalo Soc. Nat. Sci., no. E9,058); summit and<br />

anal areas, X11.7. [Explanation.—An, anal opening; CR,<br />

cryptodeltoid; HD, hypodeltoid; 0, oral plate; S, spiracle;<br />

Su, superdeltoid.1

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