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ECHINODERMATA - KU ScholarWorks - University of Kansas

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76 THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS<br />

gins. Radials five, each hexagonal in side view, 5 mm.<br />

long by 3.5 mm. wide, with shallow sinus 4 mm. long by<br />

1.5 mm. wide, with growth striae subparallel to margins.<br />

Deltoids overlap radials.<br />

Deltoids four, lancet-shaped, each 3 mm. long by 2 mm.<br />

wide, notched at the adorai tip by two spiracles along<br />

ambulacral margins. On the anal side there are ?two<br />

deltoids; a horseshoe-shaped or V-shaped ?epideltoid<br />

adjacent to the oral opening, curving around the adorai<br />

part <strong>of</strong> the anal opening on three sides, and a large<br />

lancet-shaped hypodeltoid on the aboral side <strong>of</strong> the anal<br />

opening, extended ventrally into a hood or coronal process.<br />

Two cryptodeltoids may be present. The anal<br />

spiracles or hydrospire canals are separated internally<br />

from the anal opening by deltoid septa, but externally<br />

only one anispiracle is visible. Thus there are nine openings<br />

around the oral opening, consisting <strong>of</strong> eight spiracles<br />

and the anispiracle. There are three hydrospire folds on<br />

each side <strong>of</strong> an ambulacrum.<br />

Ambulacra five, broadly linear, each 7.5 mm. long<br />

by 1.5 mm. wide, each lancet plate being exposed onethird<br />

or more <strong>of</strong> its width along the middle <strong>of</strong> each<br />

ambulacrum. The side plates are normally disposed, with<br />

30 side plates in a space <strong>of</strong> 10 mm. along an ambulacrum,<br />

one pore between adjacent side plates notched in the<br />

deltoid and radial ambulacral margins, and three main<br />

cover-plate sockets to each side plate along the main food<br />

groove. In addition to fine growth striae, the calyx plates<br />

are ornamented by coarse ridges subparallel to the plate<br />

margins.<br />

Remarks.—The type specimens are grouped together<br />

in two glass vials and it has not been possible to find all<br />

<strong>of</strong> the figured specimens. There are 37 specimens in all,<br />

including the holotype; the above description applies to<br />

specimen RX-134h. The other specimens resemble this<br />

one closely and all belong to the same species.<br />

Occurrence.—Lower Mississippian, lower Burlington<br />

Limestone, in chert, Louisiana, Missouri.<br />

Types.—Holotype, RX-134; paratypes, RX-134a-i (36<br />

specimens). Holotype unfigured but almost same as<br />

134h, herein figured on pl. 45, figs. 1-3. RX-134a, Row-<br />

LEY (1901), pl. 28, fig. 8, herein figured on pl. 45, figs.<br />

4, 6, 8. RX-134b, polished section showing hydrospires.<br />

RX-134c shows area <strong>of</strong> stem attachment. RX-134d, Row-<br />

LEY & HARE (1891), pl. 3, fig. 17, refigured by ROWLEY<br />

(1901), pl. 28, fig. 5, herein figured on pl. 45, fig. 7.<br />

RX-134e, ROWLEY (1900), pl. 2, fig. 18. RX-134f, Row-<br />

LEY (1901), pl. 28, figs. 6-7. RX-134g, 18 unfigured types.<br />

RX-134h was provisionally selected as lectotype until recently<br />

when the holotype was found. RX-134i, 11 specimens<br />

in a glass vial, with the holotype. Rowley collection,<br />

Department <strong>of</strong> Geology, <strong>University</strong> <strong>of</strong> Illinois.<br />

Genus MESOBLASTUS Etheridge & Carpenter, 1886<br />

Type-species, by original designation.—Pentatrematites crenulatus<br />

ROEMER, 1851.<br />

Generic diagnosis.—Spiraculate blastoids with nine<br />

spiracles or eight spiracles and an anispiracle, with<br />

superdeltoid, two cryptodeltoids, and hypodeltoid on<br />

the anal side, and three hydrospire folds on each side<br />

<strong>of</strong> an ambulacrum; thick hydrospire plate present; approximately<br />

five hydrospire pores to each side plate<br />

infolded into a double row; lancet covered by side<br />

plates except at adoral end; pores absent along deltoid<br />

margins; radials overlapping deltoids; calyx form subglobular.<br />

Lower Carboniferous, Belgium, England,<br />

Germany.<br />

Remarks.—The genus Mesoblastus may have been<br />

derived from a form similar to Pentremitella, but until<br />

the latter is better understood this is only a guess.<br />

Pentremitella appears to have characters similar to<br />

Schizotremites but the latter is not well understood<br />

at present.<br />

MESOBLASTUS CRENULATUS (Roemer)<br />

Plate 47, figures 1-13; text-figs. 151-157<br />

Pentatrematites crenulatus ROEMER, 1851, p. 366, pl. 4, figs. 15a-d.<br />

Description.—Calyx calcitic, subglobular in side view,<br />

rounded pentagonal in top view, 6 mm. long by 7.5 mm.<br />

wide, with flattened summit, flat to slightly convex base,<br />

and periphery below mid-height. Stem round, 0.75 mm.<br />

in diameter, on raised hexagonal area <strong>of</strong> basais, with<br />

about 30 crenellae extending almost to central round<br />

lumen. Basal circlet pentagonal in aboral view, slightly<br />

EXPLANATION OF PLATE 23<br />

FIGURE<br />

PAGE<br />

1 -3. Cordyloblastus gilbertsoni (ETHERIDGE & CARPENTER),<br />

plesiotype, 313, Schultze coll., Harvard Mus. Comp.<br />

Zoology; Middle Devonian, Pelm, Germany; oral, "D"<br />

ambulacral, aboral views <strong>of</strong> a specimen that closely resembles<br />

calyx figured by Schultze (1867, pl. 13, fig. 7c). 57<br />

4-8. Cordyloblastus acutangtdus (ScHuurzE), holotype, 325,<br />

Schultze coll. (labelled type), Harvard Mus. Comp.<br />

Zoology; Middle Devonian, Eifel, Pelm, Germany; 4-6,<br />

oral, "D" ambulacral, aboral views <strong>of</strong> a specimen exactly<br />

like type figured by Schultze (1867, pl. 13, fig. 6) ( X2.5)<br />

[another specimen in the same tray is identical with<br />

Scum..-rzE's, fig. 6c; the summit <strong>of</strong> this specimen .1 ground<br />

and polished]; 7, detail view <strong>of</strong> oral area <strong>of</strong> specimen<br />

shown in fig. 4 ( X10.5); 8, detail view <strong>of</strong> "B" ambulacrum<br />

showing pores between plates along raised ambulacral<br />

margins and lancet completely covered by side plates<br />

( X 10.6) 54

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