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ECHINODERMATA - KU ScholarWorks - University of Kansas

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BLASTOID STUDIES 61<br />

CRYPTOBLASTUS MELO (Owen & Shumard)<br />

Plate 38, figures 6-9; text-figs. 98-100<br />

Pentremites melo OWEN & SHUMARD, 1850, p. 65, pl. 7, figs. 14a-c.<br />

Description.—Calyx calcitic, subspherical, 13 mm. long<br />

by 11 mm. wide, with truncated summit and depressed<br />

base. Stem round, 1 mm. in diameter, with approximately<br />

40 crenellae extending almost one-half the radial distance<br />

in from the circumference. Basal circlet pentagonal, 3<br />

mm. wide, depressed in basal concavity, with three normally<br />

disposed basals. Radials five, elongate, recurved<br />

below, each 12 mm. long by 5.5 mm. wide, with narrow<br />

sinus and depressed interradial sutures; radials overlapping<br />

deltoids except at the adoral tip <strong>of</strong> radial limbs<br />

where in reverse manner aboral ends <strong>of</strong> the deltoid bodies<br />

slightly overlap the radial limbs, giving the appearance<br />

that the radial limbs are wedged in the deltoid bodies.<br />

Deltoids four, arrow-shaped, each 1.5 mm. long by 1.5<br />

mm. wide, with two spiracles notched in the adoral tip<br />

<strong>of</strong> each on the ambulacral margins, and without pores<br />

along margins <strong>of</strong> the ambulacra. On the anal side a large<br />

oval anispiracle occurs between an epideltoid and hypodeltoid,<br />

but internally, in a specimen seen in the Philadelphia<br />

Academy <strong>of</strong> Sciences collection, there probably<br />

are two hidden cryptodeltoids beneath the hypodeltoid.<br />

If this is true, the plate adjacent to the oral opening on<br />

the adorai side <strong>of</strong> the anal opening should probably be<br />

called the superdeltoid, instead <strong>of</strong> epideltoid.<br />

Ambulacra five, linear, each 15 mm. long by 1.25 mm.<br />

wide, recurved below, lancet exposed along the middle<br />

one-third <strong>of</strong> its width except at the aboral end. The side<br />

plates are normally disposed with 30 side plates in a<br />

space <strong>of</strong> 10 mm. length along an ambulacrum. Four coverplate<br />

sockets to each side plate are seen along the main<br />

food groove and three side cover-plate sockets to each<br />

side plate along side food grooves. Two hydrospire folds<br />

occur on each side <strong>of</strong> an ambulacrum, with a hydrospire<br />

plate on each side, and more pores than side plates (approximately<br />

two to each side plate) along the radial margins.<br />

The surfaces <strong>of</strong> the calyx plates are ornamented<br />

with fine granules aligned parallel to plate margins.<br />

The above description is based on the newly selected<br />

holotype.<br />

Remarks.—The original holotype was destroyed in a<br />

fire at Indiana <strong>University</strong> in 1883, where the Owen<br />

collection was housed. The neotype, or new holotype,<br />

here selected, is that figured by ETHERIDGE & CARPENTER,<br />

1886, pl. 7, fig. 14.<br />

Occurrence.—Lower Mississippian, lower part <strong>of</strong> Burlington<br />

Limestone, Burlington, Iowa (neotype).<br />

Type.—Neotype (new holotype), S4,959, one specimen,<br />

old number S3,766, Springer collection, U. S. National<br />

Museum, Washington. This is also the type-species<br />

<strong>of</strong> Cry ptoblastus.<br />

CRYPTOBLASTUS SHUMARDI (Meek &Worthen)<br />

Plate 38, figure 4<br />

Granatocrinus shumardi MEEK & WORTHEN, 1866, p. 257.<br />

Description.—The specimen is 36 mm. long by 17<br />

mm. wide and is here illustrated to show the shape <strong>of</strong> the<br />

specimen with brachioles attached.<br />

Occurrence.—Lower Mississippian, lower part <strong>of</strong> Burlington<br />

Limestone, Burlington, Iowa.<br />

Types.—Topotypes, S3,765, 10 specimens, Springer<br />

collection, U. S. National Museum, Washington.<br />

Genus DEVONOBLASTUS Reimann, 1935<br />

Type-species, by subsequent designation (Reimann, 1942) —l'entremites<br />

leda HALL, 1862.<br />

Generic diagnosis.—Spiraculate blastoids with five<br />

spiracles, or four spiracles in addition to an anispiracle,<br />

with anispiracle between a superdeltoid and hyperdeltoid<br />

and with two hidden cryptodeltoids, approximately<br />

five hydrospire folds on each side <strong>of</strong> an ambulacrum,<br />

lancet covered by side plates except at adoral end, with<br />

one pore between side plates along radial and part <strong>of</strong><br />

deltoid margins; deltoids visible in side view, overlapped<br />

by radials; calyx form elliptical, with conical<br />

pelvis. Devonian, New York, Ontario, ?Indiana.<br />

Remarks.—The genus Devon oblastus was probably<br />

derived from Hyperoblastus by complete closure <strong>of</strong><br />

the radial sinus, downward migration <strong>of</strong> the deltoid<br />

plates, and partial outward migration <strong>of</strong> the adorai<br />

end <strong>of</strong> the lancet plates.<br />

EXPLANATION OF PLATE 16<br />

FIGURE PAGE<br />

1-3. Orophocrinus gracilis (MEEK & WORTHEN), topotypes,<br />

S3,228, Springer coll., U.S. Natl. Mus.; Lower Mississippian,<br />

Lower Burlington Limestone, Burlington, Iowa; 1,<br />

oral view <strong>of</strong> specimen ( X4.4); 2,3, aboral and "D" ambulacral<br />

views <strong>of</strong> another specimen ( X3.6) 37<br />

4-6. Orophocrinus orbignyanus (DE KoNnvcx), topotype,<br />

S3,229, Springer coll., U.S. Natl. Mus.; Lower Carboniferous<br />

limestone, Tournai, Belgium; 4, oral view, ( X3.5);<br />

5, "B" ambulacral view, ( X2.8); 6, aboral view, ( X3.7) 37<br />

7-9. Orophocrinus verus (CUMBERLAND), plesiotype, S3,237,<br />

Springer coll., U.S. Natl. Mus.; Lower Carboniferous limestone,<br />

Lancashire, England; oral, "D" ambulacral, aboral<br />

views (all X3.5) 39<br />

10-12. Orophocrinus puzos (MONSTER), topotype, S3,230,<br />

Springer coll., U.S. Natl. Mus.; Lower Carboniferous limestone,<br />

Tournai, Belgium; oral, "D" ambulacral, aboral<br />

views (all x6.6) 38

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