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ECHINODERMATA - KU ScholarWorks - University of Kansas

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BLASTOID STUDIES 49<br />

Deltoids four, arrow-shaped, with moderately high<br />

coronal processes, and seven or eight exposed hydrospire<br />

slits on each side <strong>of</strong> an ambulacrum, extending to adjacent<br />

radial limbs. On the anal side there are three deltoid<br />

plates. The hexagonal superdeltoid is adjacent to the<br />

oral opening and the horseshoe-shaped subdeltoid rests on<br />

the inner aboral surface <strong>of</strong> the superdeltoid. The subdeltoid<br />

limbs curve around the sides <strong>of</strong> the anal opening and<br />

meet the truncated radial limbs. It is assumed that a<br />

hypodeltoid plate was present on the aboral side <strong>of</strong> the<br />

anal opening, where a facet is present. Hydrospire slits<br />

are lacking on anal side; thus there are eight exposed<br />

hydrospire fields.<br />

Ambulacra five, linear, each 3.5 mm. long by 0.5 mm.<br />

wide, with approximately 50 side plates in 10 mm. The<br />

primary and secondary side plates are pentagonal and<br />

almost equal in size, with four or five cover-plate sockets<br />

to each side plate along the main food groove. The surfaces<br />

<strong>of</strong> the calyx plates are ornamented with fine growth<br />

lines parallel to plate margins.<br />

Occurrence.—Middle Devonian, Partridge Point Formation,<br />

Partridge Point, Michigan.<br />

Types.—Plesiotypes, E21,142, 12 specimens, Reimann<br />

collection, Buffalo Society <strong>of</strong> Natural Sciences, Buffalo,<br />

New York.<br />

TRIONOBLASTUS PYRAMIDATUS (Shumard)<br />

Plate 4, figures 7-9; Plate 12, figure 8; text-figs. 67, 68<br />

Codaster pyramidatus SHUMARD, 1858, p. 238, pl. 9, figs. la-c.<br />

Description.—Calyx silicified, obconical in side view,<br />

pentagonal in oral view, 12 mm. long by 8 mm. wide,<br />

with vault 2 mm. long, pelvis 10 mm. long, pelvic angle<br />

45 degrees, and periphery at radial lips well above midheight.<br />

Basal circlet rounded in aboral view, broadly triangular<br />

near area <strong>of</strong> stem attachment, 3.5 mm. long by<br />

4 mm. wide, with three normally disposed basals. Stem<br />

round, 1 mm. in diameter, with small round lumen.<br />

Radials five, each broadly pentagonal in side view, 8 mm.<br />

long by 4 mm. wide, with broad flat sinus 3 mm. long<br />

by 3.5 mm. wide, and low limbs. The interradial sutures<br />

are in depressed areas flanked by a double or triple row<br />

<strong>of</strong> parallel ridges on either side; radials overlapping deltoids.<br />

Deltoids four, arrow-shaped, with low crests, and seven<br />

hydrospire slits on each side <strong>of</strong> an ambulacrum, extending<br />

to adjacent radial limbs. On the anal side there<br />

are three deltoids. The arrow-shaped superdeltoid is<br />

adjacent to the oral opening. The subdeltoid is horseshoeshaped,<br />

adjacent to the aboral surface <strong>of</strong> the superdeltoid,<br />

with limbs that extend on both sides <strong>of</strong> the anal opening<br />

and meet the adjacent radial limbs which overlap the<br />

subdeltoid limbs. It is assumed that a small hypodeltoid<br />

plate covered the aboral part <strong>of</strong> the anal opening and<br />

abutted against the radial limbs. Hydrospire slits are absent<br />

on anal side; therefore there are eight hydrospire<br />

fields.<br />

Ambulacra five, linear, each 4.5 mm. long by 1 mm.<br />

wide, with lancet covered by side plates, and 50 side<br />

plates in 10 mm. The side plates appear to be normally<br />

disposed. The surfaces <strong>of</strong> the calyx plates appear to be<br />

ornamented with fine growth lines parallel to plate margins.<br />

Remarks.—The flat summit and ridges parallel to the<br />

interradial sutures serve to characterize this species. The<br />

variously described species <strong>of</strong> Trionoblastus may readily<br />

be distinguished by the number <strong>of</strong> hydrospire slits in each<br />

field, and the relative height <strong>of</strong> the deltoid crests.<br />

Occurrence.—Middle Devonian, Onondaga (Columbus)<br />

Limestone, Columbus, Ohio.<br />

Types.—Plesiotypes, 37,759, eight specimens, U. S.<br />

National Museum, Washington; E21,071, seven specimens<br />

used for the above description, Buffalo Society <strong>of</strong><br />

Natural Sciences, Buffalo, New York.<br />

Order SPIRACULATA Jaekel, 1918<br />

Blastoids with hidden hydrospire slits that open<br />

into hydrospire canals, haying definite spiracles and<br />

hydrospire pores. Silurian -Permian, North America,<br />

Europe, Timor, Australia, Asia, North Africa.<br />

EXPLANATION OF PLATE 12<br />

FIGURE PAGE<br />

1-7,9. Codaster acutus M'Cov, plesiotype (figs. 1-3), S3,218,<br />

Springer coll., U.S. Natl. Mus., Lower Carboniferous limestone,<br />

Lancashire, England; plesiotypes (figs. 4-7,9), 367,<br />

Harvard Mus. Comp. Zoology, Lower Carboniferous limestone,<br />

Malham, Yorkshire, England; 1-3, oral, "D" ambulacral,<br />

aboral views (all X3.7); 4-6, oral, "D" ambulacral,<br />

aboral views (all X4.6); 7, enlarged view <strong>of</strong> "E"<br />

ambulacrum <strong>of</strong> another specimen ( X16); 9, thin sec. <strong>of</strong><br />

anal area <strong>of</strong> a third specimen, showing epideltoid against<br />

radial limbs (anal area in white) ( X16) 23<br />

8. Trionoblastus pyramidatus (SHumARD), plesiotype, 37,759,<br />

U.S. Natl. Mus.; Devonian, Onondaga Limestone, Columbus,<br />

Ohio; oral view, for comparison with Codaster, showing<br />

subdeltoid plate on adorai side <strong>of</strong> anal opening, adjacent<br />

to superdeltoid plate, with distinct suture between<br />

the two plates ( 7) 49

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