ECHINODERMATA - KU ScholarWorks - University of Kansas
ECHINODERMATA - KU ScholarWorks - University of Kansas
ECHINODERMATA - KU ScholarWorks - University of Kansas
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BLASTOID STUDIES 49<br />
Deltoids four, arrow-shaped, with moderately high<br />
coronal processes, and seven or eight exposed hydrospire<br />
slits on each side <strong>of</strong> an ambulacrum, extending to adjacent<br />
radial limbs. On the anal side there are three deltoid<br />
plates. The hexagonal superdeltoid is adjacent to the<br />
oral opening and the horseshoe-shaped subdeltoid rests on<br />
the inner aboral surface <strong>of</strong> the superdeltoid. The subdeltoid<br />
limbs curve around the sides <strong>of</strong> the anal opening and<br />
meet the truncated radial limbs. It is assumed that a<br />
hypodeltoid plate was present on the aboral side <strong>of</strong> the<br />
anal opening, where a facet is present. Hydrospire slits<br />
are lacking on anal side; thus there are eight exposed<br />
hydrospire fields.<br />
Ambulacra five, linear, each 3.5 mm. long by 0.5 mm.<br />
wide, with approximately 50 side plates in 10 mm. The<br />
primary and secondary side plates are pentagonal and<br />
almost equal in size, with four or five cover-plate sockets<br />
to each side plate along the main food groove. The surfaces<br />
<strong>of</strong> the calyx plates are ornamented with fine growth<br />
lines parallel to plate margins.<br />
Occurrence.—Middle Devonian, Partridge Point Formation,<br />
Partridge Point, Michigan.<br />
Types.—Plesiotypes, E21,142, 12 specimens, Reimann<br />
collection, Buffalo Society <strong>of</strong> Natural Sciences, Buffalo,<br />
New York.<br />
TRIONOBLASTUS PYRAMIDATUS (Shumard)<br />
Plate 4, figures 7-9; Plate 12, figure 8; text-figs. 67, 68<br />
Codaster pyramidatus SHUMARD, 1858, p. 238, pl. 9, figs. la-c.<br />
Description.—Calyx silicified, obconical in side view,<br />
pentagonal in oral view, 12 mm. long by 8 mm. wide,<br />
with vault 2 mm. long, pelvis 10 mm. long, pelvic angle<br />
45 degrees, and periphery at radial lips well above midheight.<br />
Basal circlet rounded in aboral view, broadly triangular<br />
near area <strong>of</strong> stem attachment, 3.5 mm. long by<br />
4 mm. wide, with three normally disposed basals. Stem<br />
round, 1 mm. in diameter, with small round lumen.<br />
Radials five, each broadly pentagonal in side view, 8 mm.<br />
long by 4 mm. wide, with broad flat sinus 3 mm. long<br />
by 3.5 mm. wide, and low limbs. The interradial sutures<br />
are in depressed areas flanked by a double or triple row<br />
<strong>of</strong> parallel ridges on either side; radials overlapping deltoids.<br />
Deltoids four, arrow-shaped, with low crests, and seven<br />
hydrospire slits on each side <strong>of</strong> an ambulacrum, extending<br />
to adjacent radial limbs. On the anal side there<br />
are three deltoids. The arrow-shaped superdeltoid is<br />
adjacent to the oral opening. The subdeltoid is horseshoeshaped,<br />
adjacent to the aboral surface <strong>of</strong> the superdeltoid,<br />
with limbs that extend on both sides <strong>of</strong> the anal opening<br />
and meet the adjacent radial limbs which overlap the<br />
subdeltoid limbs. It is assumed that a small hypodeltoid<br />
plate covered the aboral part <strong>of</strong> the anal opening and<br />
abutted against the radial limbs. Hydrospire slits are absent<br />
on anal side; therefore there are eight hydrospire<br />
fields.<br />
Ambulacra five, linear, each 4.5 mm. long by 1 mm.<br />
wide, with lancet covered by side plates, and 50 side<br />
plates in 10 mm. The side plates appear to be normally<br />
disposed. The surfaces <strong>of</strong> the calyx plates appear to be<br />
ornamented with fine growth lines parallel to plate margins.<br />
Remarks.—The flat summit and ridges parallel to the<br />
interradial sutures serve to characterize this species. The<br />
variously described species <strong>of</strong> Trionoblastus may readily<br />
be distinguished by the number <strong>of</strong> hydrospire slits in each<br />
field, and the relative height <strong>of</strong> the deltoid crests.<br />
Occurrence.—Middle Devonian, Onondaga (Columbus)<br />
Limestone, Columbus, Ohio.<br />
Types.—Plesiotypes, 37,759, eight specimens, U. S.<br />
National Museum, Washington; E21,071, seven specimens<br />
used for the above description, Buffalo Society <strong>of</strong><br />
Natural Sciences, Buffalo, New York.<br />
Order SPIRACULATA Jaekel, 1918<br />
Blastoids with hidden hydrospire slits that open<br />
into hydrospire canals, haying definite spiracles and<br />
hydrospire pores. Silurian -Permian, North America,<br />
Europe, Timor, Australia, Asia, North Africa.<br />
EXPLANATION OF PLATE 12<br />
FIGURE PAGE<br />
1-7,9. Codaster acutus M'Cov, plesiotype (figs. 1-3), S3,218,<br />
Springer coll., U.S. Natl. Mus., Lower Carboniferous limestone,<br />
Lancashire, England; plesiotypes (figs. 4-7,9), 367,<br />
Harvard Mus. Comp. Zoology, Lower Carboniferous limestone,<br />
Malham, Yorkshire, England; 1-3, oral, "D" ambulacral,<br />
aboral views (all X3.7); 4-6, oral, "D" ambulacral,<br />
aboral views (all X4.6); 7, enlarged view <strong>of</strong> "E"<br />
ambulacrum <strong>of</strong> another specimen ( X16); 9, thin sec. <strong>of</strong><br />
anal area <strong>of</strong> a third specimen, showing epideltoid against<br />
radial limbs (anal area in white) ( X16) 23<br />
8. Trionoblastus pyramidatus (SHumARD), plesiotype, 37,759,<br />
U.S. Natl. Mus.; Devonian, Onondaga Limestone, Columbus,<br />
Ohio; oral view, for comparison with Codaster, showing<br />
subdeltoid plate on adorai side <strong>of</strong> anal opening, adjacent<br />
to superdeltoid plate, with distinct suture between<br />
the two plates ( 7) 49