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ECHINODERMATA - KU ScholarWorks - University of Kansas

ECHINODERMATA - KU ScholarWorks - University of Kansas

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1 1 0 THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS<br />

new genus; so-called Metablastus hispanicus <strong>of</strong> the<br />

Devonian <strong>of</strong> Spain, probably does not belong to this<br />

genus; forms identified as Orbitremites aplatus, O.<br />

calycinus, O. excavatus, O. fimbriatus, O. grandis, O.<br />

magnificus, O. ornatus, O. parvus, O. spathatus, O.<br />

spinuliferus, and O. tvachsmuthi probably belong to<br />

diverse genera, some perhaps to new genera, but seemingly<br />

not to Orbitremites; Phaenoschisma? kentuckyense<br />

probably does not belong to Phaenoschisrna. It<br />

is hoped that others will restudy blastoids in collections<br />

<strong>of</strong> various museums in order to fill in gaps<br />

in knowledge such as are here suggested.<br />

It is significant that very few blastoids have been<br />

described from Pennsylvanian deposits and that records<br />

<strong>of</strong> blastoids in South America, Africa, Antarctica,<br />

Asia, and the North Polar regions are meager or lacking.<br />

Evidently the study <strong>of</strong> blastoids is not yet far<br />

advanced. It is premature to say that the center <strong>of</strong><br />

dispersal <strong>of</strong> blastoids is the mid-Continent region <strong>of</strong><br />

the United States, merely because the oldest and most<br />

primitive blastoids occur in Silurian deposits <strong>of</strong> this<br />

region. The basis for this sort <strong>of</strong> statement is that<br />

Silurian blastoids have not been described from other<br />

continents, but it would not be surprising if they are<br />

discovered in various regions outside <strong>of</strong> North<br />

America.<br />

There is little in common between blastoid genera<br />

<strong>of</strong> Europe and North America, or between those <strong>of</strong><br />

Timor and Europe, or between Timor and North<br />

America, and yet when lineages are traced, a tie between<br />

these regions seems to exist. "Missing links,"<br />

both stratigraphic and geographic, are postulated to<br />

account for the differences; when missing links are<br />

found, the literature on blastoids will probably be<br />

double its present size. To date, approximately 800<br />

articles have been published on blastoids; they treat<br />

some 33 genera <strong>of</strong> the Fissiculata and 35 genera <strong>of</strong> the<br />

Spiraculata, together including about 350 species.<br />

KEY TO FAMILIES AND GENERA OF<br />

BLASTOIDEA<br />

The following key is <strong>of</strong>fered as an aid to those<br />

who may wish to identify genera <strong>of</strong> blastoids, as now<br />

known and understood. The key is arranged by families<br />

belonging to the two orders Fissiculata and<br />

Spiraculata. In using the key, one should proceed by<br />

noting the characters recorded under succesive letters<br />

and numbers until the desired end is reached.<br />

Hydrospire fields exposed or spiracular slits<br />

present<br />

FISSICULATA<br />

Hydrospire fields concealed, pores and<br />

spiracles present<br />

SPIRACULATA<br />

FISSICULATA [Families, see nos. 1-5]<br />

1. Exposed hydrospire fields ten (Phaenoschismatidae)<br />

A<br />

2. Exposed hydrospire fields nine<br />

(Brachyschismatidae)<br />

3. Exposed hydrospire fields eight (Codasteridae)<br />

4. Spiracular slits ten (Orophocrinidae) D<br />

5. Spiracular slits eight (Astrocrinidae)<br />

SPIRACULATA [Families, see nos. 6-10]<br />

6. Spiracles five, simple (Pentremitidae)<br />

7. Spiracles five, paired (Troosticrinidae) G<br />

8. Spiracles nine (Granatocrinidae) H<br />

9. Spiracles ten, anus separate, cryptodeltoids<br />

exposed (Nucleocrinidae)<br />

10. Spiracles ten, anus separate, cryptodeltoids<br />

absent (Schizoblastidae)<br />

A. PHAENOSCHISMATIDAE (Silurian-Permian)<br />

[Genera, see nos. 11 -311<br />

11. Anal deltoids four to six 14, 15<br />

12. Anal deltoids three Decaschisma<br />

13. Anal deltoids two or one 16, 17<br />

14. Hydrospire slits reduced in number on<br />

anal side<br />

Deltoschisma<br />

15. Hydrospire slits not reduced in number<br />

on anal side 18, 19<br />

16. Ambulacra with lancets widely exposed<br />

Phaenoblastus<br />

17. Ambulacra with lancets covered by<br />

side plates 22, 23<br />

18. Ambulacra directed laterally outward,<br />

six anal deltoids<br />

Polydeltoideus<br />

19. Ambulacra directed downward, four<br />

anal deltoids 20, 21<br />

20. Hydrospire slits completely exposed .. Pleuroschisma<br />

21. Hydrospire slits almost hidden, with<br />

formation <strong>of</strong> pores<br />

Hyperoblastus<br />

22. Ambulacra five 24, 25<br />

23. Ambulacra two Dipteroblastus<br />

24. Ambulacra extremely long 26, 27<br />

25. Ambulacra moderately long to extremely short 28, 29<br />

26. Ambulacra flaring outward into elongate<br />

wings<br />

Thaumatoblastus<br />

27. Ambulacra projecting downward aborally,<br />

calyx globular Nymphaeoblastus<br />

28. Ambulacra moderately long 30a, 30b, 31<br />

29. Ambulacra extremely short Neoschisma<br />

30a.Calyx conical, ambulacra narrow Con oschisma<br />

30b. Calyx conical, ambulacra broad Phaenoschisma<br />

31. Calyx flattened Notoblastus

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