ECHINODERMATA - KU ScholarWorks - University of Kansas
ECHINODERMATA - KU ScholarWorks - University of Kansas
ECHINODERMATA - KU ScholarWorks - University of Kansas
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1 1 0 THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS<br />
new genus; so-called Metablastus hispanicus <strong>of</strong> the<br />
Devonian <strong>of</strong> Spain, probably does not belong to this<br />
genus; forms identified as Orbitremites aplatus, O.<br />
calycinus, O. excavatus, O. fimbriatus, O. grandis, O.<br />
magnificus, O. ornatus, O. parvus, O. spathatus, O.<br />
spinuliferus, and O. tvachsmuthi probably belong to<br />
diverse genera, some perhaps to new genera, but seemingly<br />
not to Orbitremites; Phaenoschisma? kentuckyense<br />
probably does not belong to Phaenoschisrna. It<br />
is hoped that others will restudy blastoids in collections<br />
<strong>of</strong> various museums in order to fill in gaps<br />
in knowledge such as are here suggested.<br />
It is significant that very few blastoids have been<br />
described from Pennsylvanian deposits and that records<br />
<strong>of</strong> blastoids in South America, Africa, Antarctica,<br />
Asia, and the North Polar regions are meager or lacking.<br />
Evidently the study <strong>of</strong> blastoids is not yet far<br />
advanced. It is premature to say that the center <strong>of</strong><br />
dispersal <strong>of</strong> blastoids is the mid-Continent region <strong>of</strong><br />
the United States, merely because the oldest and most<br />
primitive blastoids occur in Silurian deposits <strong>of</strong> this<br />
region. The basis for this sort <strong>of</strong> statement is that<br />
Silurian blastoids have not been described from other<br />
continents, but it would not be surprising if they are<br />
discovered in various regions outside <strong>of</strong> North<br />
America.<br />
There is little in common between blastoid genera<br />
<strong>of</strong> Europe and North America, or between those <strong>of</strong><br />
Timor and Europe, or between Timor and North<br />
America, and yet when lineages are traced, a tie between<br />
these regions seems to exist. "Missing links,"<br />
both stratigraphic and geographic, are postulated to<br />
account for the differences; when missing links are<br />
found, the literature on blastoids will probably be<br />
double its present size. To date, approximately 800<br />
articles have been published on blastoids; they treat<br />
some 33 genera <strong>of</strong> the Fissiculata and 35 genera <strong>of</strong> the<br />
Spiraculata, together including about 350 species.<br />
KEY TO FAMILIES AND GENERA OF<br />
BLASTOIDEA<br />
The following key is <strong>of</strong>fered as an aid to those<br />
who may wish to identify genera <strong>of</strong> blastoids, as now<br />
known and understood. The key is arranged by families<br />
belonging to the two orders Fissiculata and<br />
Spiraculata. In using the key, one should proceed by<br />
noting the characters recorded under succesive letters<br />
and numbers until the desired end is reached.<br />
Hydrospire fields exposed or spiracular slits<br />
present<br />
FISSICULATA<br />
Hydrospire fields concealed, pores and<br />
spiracles present<br />
SPIRACULATA<br />
FISSICULATA [Families, see nos. 1-5]<br />
1. Exposed hydrospire fields ten (Phaenoschismatidae)<br />
A<br />
2. Exposed hydrospire fields nine<br />
(Brachyschismatidae)<br />
3. Exposed hydrospire fields eight (Codasteridae)<br />
4. Spiracular slits ten (Orophocrinidae) D<br />
5. Spiracular slits eight (Astrocrinidae)<br />
SPIRACULATA [Families, see nos. 6-10]<br />
6. Spiracles five, simple (Pentremitidae)<br />
7. Spiracles five, paired (Troosticrinidae) G<br />
8. Spiracles nine (Granatocrinidae) H<br />
9. Spiracles ten, anus separate, cryptodeltoids<br />
exposed (Nucleocrinidae)<br />
10. Spiracles ten, anus separate, cryptodeltoids<br />
absent (Schizoblastidae)<br />
A. PHAENOSCHISMATIDAE (Silurian-Permian)<br />
[Genera, see nos. 11 -311<br />
11. Anal deltoids four to six 14, 15<br />
12. Anal deltoids three Decaschisma<br />
13. Anal deltoids two or one 16, 17<br />
14. Hydrospire slits reduced in number on<br />
anal side<br />
Deltoschisma<br />
15. Hydrospire slits not reduced in number<br />
on anal side 18, 19<br />
16. Ambulacra with lancets widely exposed<br />
Phaenoblastus<br />
17. Ambulacra with lancets covered by<br />
side plates 22, 23<br />
18. Ambulacra directed laterally outward,<br />
six anal deltoids<br />
Polydeltoideus<br />
19. Ambulacra directed downward, four<br />
anal deltoids 20, 21<br />
20. Hydrospire slits completely exposed .. Pleuroschisma<br />
21. Hydrospire slits almost hidden, with<br />
formation <strong>of</strong> pores<br />
Hyperoblastus<br />
22. Ambulacra five 24, 25<br />
23. Ambulacra two Dipteroblastus<br />
24. Ambulacra extremely long 26, 27<br />
25. Ambulacra moderately long to extremely short 28, 29<br />
26. Ambulacra flaring outward into elongate<br />
wings<br />
Thaumatoblastus<br />
27. Ambulacra projecting downward aborally,<br />
calyx globular Nymphaeoblastus<br />
28. Ambulacra moderately long 30a, 30b, 31<br />
29. Ambulacra extremely short Neoschisma<br />
30a.Calyx conical, ambulacra narrow Con oschisma<br />
30b. Calyx conical, ambulacra broad Phaenoschisma<br />
31. Calyx flattened Notoblastus