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ECHINODERMATA - KU ScholarWorks - University of Kansas

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82 THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS<br />

aboral side <strong>of</strong> the anal opening and abuts against the adjacent<br />

radial limbs. The cryptodeltoids are partially exposed<br />

externally and therefore a paired anispiracle occurs<br />

on the anal side. Thus, one paired anispiracle and four<br />

paired spiracles are recognized, a total <strong>of</strong> five paired<br />

spiracles around the oral opening.<br />

Ambulacra five, short, linear, each 15 mm. long by<br />

1.5 mm. wide, with lancet covered by side plates, and 22<br />

side plates in a space <strong>of</strong> 10 mm. along an ambulacrum.<br />

Side plates poorly preserved but appearing to be normally<br />

disposed. The surfaces <strong>of</strong> calyx plates are worn but the<br />

ornamentation appears to consist <strong>of</strong> fine growth lines subparallel<br />

to plate margins.<br />

Occurrence.—Upper Mississippian, Salem Limestone,<br />

Ellettsville, Indiana.<br />

Type.—Holotype, 6,323, one specimen, Gurley collection,<br />

Walker Museum, <strong>University</strong> <strong>of</strong> Chicago.<br />

METABLASTUS WORTHENI (Hall)<br />

Plate 50, figures 4-9; text-figs. 161-163<br />

Pentremites wortheni HALL, 1858, p. 606, pl. 15, fig. 1.<br />

Description.—Calyx calcitic, 16 mm. long by 7.5 mm.<br />

wide, bipyramidal in side view, with flat summit and flat<br />

triangular base, pentagonal in oral view, with periphery<br />

well above mid-height at radial lips. Vault 6.5 mm.<br />

long, pelvis 9.5 mm. long, and pelvic angle 30 degrees.<br />

Stem impression appears to be round, about 1 mm. in<br />

diameter, crenellar, with small round lumen, resting on<br />

the flat triangular aboral surface. Basal circlet long,<br />

rounded pentagonal in basal view, 5 mm. long by 5 mm.<br />

wide, triangular at base, with three normally disposed<br />

basals, each <strong>of</strong> which has a high, wide ridge extending<br />

down the mid-line from the aboral end. The ridge on<br />

the azygous basal bifurcates at half the distance upward<br />

adorally, each resulting ridge extending to the<br />

radial lips <strong>of</strong> the "A" and "B" ambulacra, respectively.<br />

The ridge on each <strong>of</strong> the other two basais trifurcates, with<br />

the median ridge <strong>of</strong> each extending to the radial lips <strong>of</strong><br />

the "C" and "E" ambulacra, respectively, and the lateral<br />

ridges extending to the lips <strong>of</strong> the "A," "B," and "D"<br />

ambulacra, respectively. These broad ridges give a bulge<br />

or flare to the pelvis in the region <strong>of</strong> the basiradial sutures,<br />

and are prevalent in the specimens <strong>of</strong> various species<br />

<strong>of</strong> Metablastus that occur in the Keokuk, Warsaw,<br />

and Salem formations. Radials five, each 10 mm. long<br />

by 3.5 mm. wide, broadly quadrangular in side view,<br />

with narrow, deep sinus 5 mm. long by 2 mm. wide;<br />

radials overlapping deltoids.<br />

Deltoids four, short, not visible in side view, each<br />

notched at the adorai end by a V-shaped paired spiracle,<br />

with deltoid lips and lancet stipes adjacent to oral opening.<br />

On the anal side there are four deltoids—a superdeltoid,<br />

two cryptodeltoids, and a hypodeltoid. The superdeltoid<br />

is adjacent to the oral opening, with two internal<br />

septa on its aboral surface separating the hydrospire canals<br />

or anal spiracles from the anal opening. Resting on these<br />

two septa are the two cryptodeltoids, which are overlapped<br />

by the adorai ends <strong>of</strong> the adjacent radial limbs.<br />

The pentagonal hypodeltoid abuts against the radial<br />

limbs, covers the cryptodeltoids, and covers the aboral part<br />

<strong>of</strong> the anal opening. The cryptodeltoids are partially exposed<br />

and therefore a paired anispiracle is formed on the<br />

anal side. Thus, the oral opening is surrounded by five<br />

openings, consisting <strong>of</strong> four paired spiracles and an anal<br />

paired spiracle or paired anispiracle.<br />

Ambulacra five, linear, each 7 mm. long by 1 mm.<br />

wide, with lancet completely covered by side plates, one<br />

pore between side plates along deltoid and radial margins,<br />

and 34 side plates in a space <strong>of</strong> 10 mm. along an ambulacrum.<br />

Side plates normally disposed. There are four or<br />

five hydrospire folds on each side <strong>of</strong> an ambulacrum. The<br />

surfaces <strong>of</strong> the calyx plates are ornamented by fine growth<br />

lines subparallel to plate margins .<br />

Occurrence.—Upper Mississippian, Warsaw Limestone,<br />

Boonville, Missouri.<br />

Types.—Plesiotypes, four specimens, purchased from<br />

Wards 6-21-41, Buffalo Society <strong>of</strong> Natural Sciences, Buffalo,<br />

New York.<br />

Genus MONADOBLASTUS Fay, n. gen.<br />

Type-species, by original designation.—Monadoblastos granulosus<br />

FAY, n. sp.<br />

Generic diagnosis.—Spiraculate blastoids with nine<br />

spiracles, or eight spiracles and an anispiracle, with<br />

anispiracle between epideltoid and hypodeltoid plates,<br />

with hydrospire plates bearing approximately two<br />

pores to each side plate along radial margins, pores<br />

absent along deltoids, lancet covered by side plates<br />

except near adorai end, one hydrospire fold on each<br />

side <strong>of</strong> an ambulacrum, radials overlapping deltoids,<br />

base concave, and form <strong>of</strong> calyx elliptical. Mississippian,<br />

New Mexico.<br />

Remarks.—The genus Monadoblastus was probably<br />

derived from Tanaoblastus by reduction <strong>of</strong> the<br />

hydrospires to one and invagination <strong>of</strong> the basal circlet.<br />

MONADOBLASTUS GRANULOSUS Fay, n. sp.<br />

Plate 37, figures 1-3, 10; text-figs. 164-167<br />

Description.—Calyx calcitic, suboval in side view, subpentagonal<br />

in oral view, 5 mm. long by 4 mm. wide,<br />

with flat summit, concave base, and periphery at midheight.<br />

Stem round, about 0.5 mm. in diameter, impression<br />

poorly preserved. Basal circlet pentagonal in<br />

aboral view, 1.25 mm. in diameter, with three normally<br />

disposed basal plates. Radials five, each broadly pentagonal<br />

in side view, recurved in basal concavity, 4 mm.<br />

long by 2.5 mm. wide, with radial sinus 4 mm. long by

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