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ECHINODERMATA - KU ScholarWorks - University of Kansas

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BLASTOID STUDIES 97<br />

Occurrence.—Middle Devonian, Ludlowville Formation<br />

(Tichenor Limestone), Cazenovia Creek, Springbrook,<br />

New York.<br />

Type.—Holotype, E11,811, one specimen, Fred Wattles<br />

collection, Buffalo Society <strong>of</strong> Natural Sciences, Buffalo,<br />

New York.<br />

PLACOBLASTUS LUCINA (Hall)<br />

Plate 32, figures 4-7, 10; text-figs. 200, 201<br />

Nucleoc-rinus lucina HALL, 1862, p. 148, pl. 1, fig. 16.<br />

Elaeacrinus lucina SHUMARD, 1866, p. 369.<br />

Olivanites lucina HAMBACH, 1903, p. 50.<br />

Description.—Calyx calcitic, flattened spherical in side<br />

view, pentagonal in top view with moderately sharp<br />

ambulacra and concave interambulacra, 9 mm. long by<br />

9 mm. wide, with slightly concave base. Stem round, 0.25<br />

mm. wide, in flat basal circlet, crenellar, with small round<br />

lumen. Basal circlet rounded pentagonal in aboral view,<br />

2.5 mm. in diameter, almost flat to slightly concave, with<br />

three normally disposed basals. Radials five, short, wide,<br />

pentagonal in side view, each 3 mm. long by 4 mm. wide,<br />

with short, shallow sinus 1.5 mm. long by 1 mm. wide,<br />

recurved below in slightly concave aboral surface. Deltoids<br />

overlap radials.<br />

Deltoids four, sublenticular, each 8 mm. long by 5<br />

mm. wide, with two hemielliptical spiracles in the adorai<br />

tip along ambulacral margins. In top view the deltoids<br />

are slightly concave inward, the aboral end <strong>of</strong> each abutting<br />

against a large arrow-shaped oral plate. On the anal<br />

side at least three anal deltoids occur, the two lenticular<br />

exposed cryptodeltoids along ambulacral margins, with<br />

the elongate subquadrangular hypodeltoid between. On<br />

the adoral side <strong>of</strong> the anal opening there are two polygonal<br />

plates between the adorai tips <strong>of</strong> the cryptodeltoids.<br />

The plate adjacent to the anal opening is here interpreted<br />

to be a superdeltoid plate, and the adorally disposed<br />

plate is interpreted to be an accessory anal oral<br />

plate. Calyx plates ornamented as in Nucleocrinus. There<br />

are two semielliptical slitlike spiracles on either side <strong>of</strong><br />

the anal opening, one each in the adorai tip <strong>of</strong> each cryptodeltoid.<br />

Thus, 11 openings surround the oral aperture,<br />

consisting <strong>of</strong> ten spiracles and the separate anal opening.<br />

There are two hydrospire folds on each side <strong>of</strong> an ambulacrum,<br />

with a moderately long hydrospire canal.<br />

Ambulacra five, linear, recurved below, each 11 mm.<br />

long by 1 mm. wide, with lancet covered by side plates,<br />

and 40 side plates in a space <strong>of</strong> 10 mm. along an ambulacrum.<br />

Side plates normally disposed, with an inner side<br />

plate, as in Nucleocrinus.<br />

Occurrence.—Middle Devonian, Hamilton beds, Most<br />

cow, New York.<br />

Types.—The holotype is on deposit at the American<br />

Museum <strong>of</strong> Natural History, Hall collection, 5038/2, and<br />

is slightly damaged, but usable for general identification <strong>of</strong><br />

other specimens. Topotypes, 13,276, Hall collection, one<br />

specimen; 19,049, two specimens, one <strong>of</strong> which is Nucleocrinus<br />

elegans, Gurley collection, Walker Museum, <strong>University</strong><br />

<strong>of</strong> Chicago. The above description is based on<br />

specimen 19,049.<br />

Genus POROBLASTUS Fay, n. gen.<br />

Type-species, by original designation (herein).—PentretnItes ( Granatocrinus?)<br />

granulosus MEEK & WORTHEN, 1865.<br />

Generic diagnosis.—Spiraculate blastoids with nine<br />

spiracles, or eight spiracles and anispiracle, the anispiracle<br />

located between an epideltoid and hypodeltoid,<br />

radials overlapping deltoids, hydrospire plate<br />

present, one pore between side plates along deltoid<br />

margins but two pores to each side plate along radial<br />

margins, one hydrospire fold on each side <strong>of</strong> an<br />

ambulacrum, lancet covered at aboral end but gradually<br />

exposed to one-third <strong>of</strong> its width near the adoral<br />

end, base concave, and shape ellipsoidal. Mississippian,<br />

Illinois, Iowa, Missouri.<br />

Remarks.—The genus Poroblastus may have been<br />

derived from Lophoblastus by infolding <strong>of</strong> radial mar-<br />

FIGURE<br />

EXPLANATION OF PLATE 32<br />

PAGE<br />

1-3. Placoblastus eriensis (REImANN), holotype, El 1,811, Fred<br />

Wattles coll., Buffalo Soc. Nat. Sci.; Middle Devonian, 8.<br />

Tichenor Limestone, Cazenovia Creek, Springbrook, N.Y.;<br />

oral, "D" ambulacral, aboral views <strong>of</strong> specimen described<br />

as Nucleocrinus verneuili eriensis (all X2.4) 97<br />

4-7,10. Placoblastus lucina (HALL), ?topotypes, 19,049 (figs<br />

4-6,10), Gurley coll., Univ. Chicago, Devonian, Hamilton<br />

Group, Moscow, N.Y., and 13,276 (fig. 7), James Hall 9.<br />

coll., Univ. Chicago, Middle Devonian, New York; 4-6,<br />

one <strong>of</strong> two specimens labelled Nucleocrinus elegans, oral,<br />

"D" ambulacral, aboral views ( X5.1); 10, specimen illustrated<br />

in fig. 4, showing large plate between adorai tips<br />

<strong>of</strong> cryptodeltoids ( X3.6); 7. Aboral view <strong>of</strong> polished<br />

cross section showing hydrospire folds <strong>of</strong> "E" ambulacrum<br />

( X12.6) 97<br />

Placoblastus angularis (LYON), plesiotype, E9,057, Reimann<br />

coll., Buffalo Soc. Nat. Sci. (one <strong>of</strong> five specimens<br />

labelled Nucleocrinus powelli; Middle Devonian, Ludlowville<br />

Formation (Tichenor Limestone), Cazenovia Creek,<br />

Springbrook, N.Y.; aboral view <strong>of</strong> polished cross section<br />

<strong>of</strong> "E" ambulacrum showing hydrospire folds ( X 9.3) 95<br />

Nucleocrinus meloniformis (BARRIs), plcsiotype, E21,120,<br />

Charles Southworth coll., Buffalo Soc. Nat. Sci.; Middle<br />

Devonian, Hungry Hollow Formation (coral zone <strong>of</strong><br />

Widder beds), tile yard, Thedford, Ontario, polished section<br />

<strong>of</strong> "E" ambulacrum showing two folds on right originating<br />

at deltoid ( X19) 86

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