ECHINODERMATA - KU ScholarWorks - University of Kansas
ECHINODERMATA - KU ScholarWorks - University of Kansas
ECHINODERMATA - KU ScholarWorks - University of Kansas
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BLASTOID STUDIES 97<br />
Occurrence.—Middle Devonian, Ludlowville Formation<br />
(Tichenor Limestone), Cazenovia Creek, Springbrook,<br />
New York.<br />
Type.—Holotype, E11,811, one specimen, Fred Wattles<br />
collection, Buffalo Society <strong>of</strong> Natural Sciences, Buffalo,<br />
New York.<br />
PLACOBLASTUS LUCINA (Hall)<br />
Plate 32, figures 4-7, 10; text-figs. 200, 201<br />
Nucleoc-rinus lucina HALL, 1862, p. 148, pl. 1, fig. 16.<br />
Elaeacrinus lucina SHUMARD, 1866, p. 369.<br />
Olivanites lucina HAMBACH, 1903, p. 50.<br />
Description.—Calyx calcitic, flattened spherical in side<br />
view, pentagonal in top view with moderately sharp<br />
ambulacra and concave interambulacra, 9 mm. long by<br />
9 mm. wide, with slightly concave base. Stem round, 0.25<br />
mm. wide, in flat basal circlet, crenellar, with small round<br />
lumen. Basal circlet rounded pentagonal in aboral view,<br />
2.5 mm. in diameter, almost flat to slightly concave, with<br />
three normally disposed basals. Radials five, short, wide,<br />
pentagonal in side view, each 3 mm. long by 4 mm. wide,<br />
with short, shallow sinus 1.5 mm. long by 1 mm. wide,<br />
recurved below in slightly concave aboral surface. Deltoids<br />
overlap radials.<br />
Deltoids four, sublenticular, each 8 mm. long by 5<br />
mm. wide, with two hemielliptical spiracles in the adorai<br />
tip along ambulacral margins. In top view the deltoids<br />
are slightly concave inward, the aboral end <strong>of</strong> each abutting<br />
against a large arrow-shaped oral plate. On the anal<br />
side at least three anal deltoids occur, the two lenticular<br />
exposed cryptodeltoids along ambulacral margins, with<br />
the elongate subquadrangular hypodeltoid between. On<br />
the adoral side <strong>of</strong> the anal opening there are two polygonal<br />
plates between the adorai tips <strong>of</strong> the cryptodeltoids.<br />
The plate adjacent to the anal opening is here interpreted<br />
to be a superdeltoid plate, and the adorally disposed<br />
plate is interpreted to be an accessory anal oral<br />
plate. Calyx plates ornamented as in Nucleocrinus. There<br />
are two semielliptical slitlike spiracles on either side <strong>of</strong><br />
the anal opening, one each in the adorai tip <strong>of</strong> each cryptodeltoid.<br />
Thus, 11 openings surround the oral aperture,<br />
consisting <strong>of</strong> ten spiracles and the separate anal opening.<br />
There are two hydrospire folds on each side <strong>of</strong> an ambulacrum,<br />
with a moderately long hydrospire canal.<br />
Ambulacra five, linear, recurved below, each 11 mm.<br />
long by 1 mm. wide, with lancet covered by side plates,<br />
and 40 side plates in a space <strong>of</strong> 10 mm. along an ambulacrum.<br />
Side plates normally disposed, with an inner side<br />
plate, as in Nucleocrinus.<br />
Occurrence.—Middle Devonian, Hamilton beds, Most<br />
cow, New York.<br />
Types.—The holotype is on deposit at the American<br />
Museum <strong>of</strong> Natural History, Hall collection, 5038/2, and<br />
is slightly damaged, but usable for general identification <strong>of</strong><br />
other specimens. Topotypes, 13,276, Hall collection, one<br />
specimen; 19,049, two specimens, one <strong>of</strong> which is Nucleocrinus<br />
elegans, Gurley collection, Walker Museum, <strong>University</strong><br />
<strong>of</strong> Chicago. The above description is based on<br />
specimen 19,049.<br />
Genus POROBLASTUS Fay, n. gen.<br />
Type-species, by original designation (herein).—PentretnItes ( Granatocrinus?)<br />
granulosus MEEK & WORTHEN, 1865.<br />
Generic diagnosis.—Spiraculate blastoids with nine<br />
spiracles, or eight spiracles and anispiracle, the anispiracle<br />
located between an epideltoid and hypodeltoid,<br />
radials overlapping deltoids, hydrospire plate<br />
present, one pore between side plates along deltoid<br />
margins but two pores to each side plate along radial<br />
margins, one hydrospire fold on each side <strong>of</strong> an<br />
ambulacrum, lancet covered at aboral end but gradually<br />
exposed to one-third <strong>of</strong> its width near the adoral<br />
end, base concave, and shape ellipsoidal. Mississippian,<br />
Illinois, Iowa, Missouri.<br />
Remarks.—The genus Poroblastus may have been<br />
derived from Lophoblastus by infolding <strong>of</strong> radial mar-<br />
FIGURE<br />
EXPLANATION OF PLATE 32<br />
PAGE<br />
1-3. Placoblastus eriensis (REImANN), holotype, El 1,811, Fred<br />
Wattles coll., Buffalo Soc. Nat. Sci.; Middle Devonian, 8.<br />
Tichenor Limestone, Cazenovia Creek, Springbrook, N.Y.;<br />
oral, "D" ambulacral, aboral views <strong>of</strong> specimen described<br />
as Nucleocrinus verneuili eriensis (all X2.4) 97<br />
4-7,10. Placoblastus lucina (HALL), ?topotypes, 19,049 (figs<br />
4-6,10), Gurley coll., Univ. Chicago, Devonian, Hamilton<br />
Group, Moscow, N.Y., and 13,276 (fig. 7), James Hall 9.<br />
coll., Univ. Chicago, Middle Devonian, New York; 4-6,<br />
one <strong>of</strong> two specimens labelled Nucleocrinus elegans, oral,<br />
"D" ambulacral, aboral views ( X5.1); 10, specimen illustrated<br />
in fig. 4, showing large plate between adorai tips<br />
<strong>of</strong> cryptodeltoids ( X3.6); 7. Aboral view <strong>of</strong> polished<br />
cross section showing hydrospire folds <strong>of</strong> "E" ambulacrum<br />
( X12.6) 97<br />
Placoblastus angularis (LYON), plesiotype, E9,057, Reimann<br />
coll., Buffalo Soc. Nat. Sci. (one <strong>of</strong> five specimens<br />
labelled Nucleocrinus powelli; Middle Devonian, Ludlowville<br />
Formation (Tichenor Limestone), Cazenovia Creek,<br />
Springbrook, N.Y.; aboral view <strong>of</strong> polished cross section<br />
<strong>of</strong> "E" ambulacrum showing hydrospire folds ( X 9.3) 95<br />
Nucleocrinus meloniformis (BARRIs), plcsiotype, E21,120,<br />
Charles Southworth coll., Buffalo Soc. Nat. Sci.; Middle<br />
Devonian, Hungry Hollow Formation (coral zone <strong>of</strong><br />
Widder beds), tile yard, Thedford, Ontario, polished section<br />
<strong>of</strong> "E" ambulacrum showing two folds on right originating<br />
at deltoid ( X19) 86