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ECHINODERMATA - KU ScholarWorks - University of Kansas

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BLASTOID STUDIES 73<br />

Ambulacra five, linear, recurved below, each 11 mm.<br />

long by 1 mm. wide, with median one-third <strong>of</strong> width<br />

<strong>of</strong> lancet exposed and 25 side plates in 10 mm. length <strong>of</strong><br />

an ambulacrum. The side plates are normally disposed,<br />

with approximately two pores to each side plate, a hydrospire<br />

plate present, and five or six main cover-plate<br />

sockets to each side plate along the main food groove.<br />

The surfaces <strong>of</strong> the calyx plates are ornamented by fine<br />

granules that are arranged along growth lines.<br />

Occurrence.—Lower Carboniferous Limestone, Lancashire<br />

and Yorkshire, England.<br />

Types.—Plesiotype, 13,877, one specimen, Gurley collection,<br />

Walker Museum, <strong>University</strong> <strong>of</strong> Chicago. Plesiotype,<br />

241, one specimen, deKoninck collection, old no.<br />

2,330, Carboniferous Limestone, Settle, Yorkshire, England,<br />

Harvard Museum <strong>of</strong> Comparative Zoology, Cambridge,<br />

Mass.<br />

Genus GLOBOBLASTUS Hambach, 1903<br />

Type-species, by original designation (monotypy), and by subsequent<br />

designation (Bassler, 1938).—Pentremites norwoodi<br />

OWEN & SHUMARD, 1850.<br />

Generic diagnosis (emend. FAY, 1960).—Spiraculate<br />

blastoids with five spiracles, composed <strong>of</strong> four<br />

spiracles and an anispiracle, with superdeltoid, two<br />

cryptodeltoids, and a hypodeltoid on the anal side;<br />

lancet covered by side plates except near adorai end,<br />

radials overlapping deltoids, approximately two pores<br />

to each side plate along radial margins, hydrospire plate<br />

present, deltoids short; two hydrospire folds on each<br />

side <strong>of</strong> an ambulacrum; calyx subglobular in shape.<br />

Mississippian, Illinois, Iowa, Missouri.<br />

Remarks.—The genus Globoblastus was probably<br />

derived from a form similar to Devonoblastus in<br />

which a hydrospire plate and infolded pores formed<br />

along the ambulacra, the hydrospires becoming reduced<br />

in number, and form <strong>of</strong> the calyx subglobular.<br />

GLOBOBLASTUS NORWOODI (Owen & Shumard)<br />

Plate 44, figures 1-7; text-figs. 139-142<br />

Pentremites norwoodi OWEN & SHUMARD, 1850, p. 64, pl. 7, figs.<br />

13a-c.<br />

Description.—The new holotype (neotype) is ellipsoidal,<br />

15 mm. long by 12 mm. wide, with concave base.<br />

Stem round, approximately 1 mm. in diameter, with about<br />

40 crenellae extending toward center. Basal circlet small,<br />

in deep concavity, with three normally disposed basais.<br />

Radials each 15 mm. long by 7 mm. wide, recurved in<br />

basal concavity, with long narrow sinuses; radials overlapping<br />

deltoids.<br />

Deltoids four, short, each 2 mm. long by 2 mm. wide,<br />

with a single spiracle at their adorai tips. On the anal<br />

side the superdeltoid is 1 mm. long by 2 mm. wide, and<br />

hypodeltoid 1 mm. long by 2 mm. wide, with anispiracle<br />

between. The two cryptodeltoids are covered over by the<br />

hypodeltoid, are infolded internally into hydrospires, and<br />

are overlapped by the radial limbs. Adorally the cryptodeltoids<br />

separate the anal opening from the adjacent hydrospire<br />

canals internally, and rest on the aboral surface<br />

<strong>of</strong> the superdeltoid. There are two hydrospire folds on<br />

each side <strong>of</strong> an ambulacrum, with a hydrospire plate<br />

present; approximately two pores to each side plate are<br />

excavated in the ambulacral margins <strong>of</strong> the radial plates.<br />

The oral opening is surrounded by the four deltoid lips<br />

and the superdeltoid, and by five spiracles, or four spiracles<br />

and an anispiracle.<br />

Ambulacra five, linear, each 16 mm. long by 1.5 mm.<br />

wide, recurved below, with lancet covered by side plates<br />

along the aboral one-half <strong>of</strong> each ambulacrum but exposed<br />

one-third <strong>of</strong> its width near the oral opening. There<br />

are 30 side plates in a space <strong>of</strong> 10 mm. along an ambulacrum,<br />

with approximately two pores to each side plate<br />

(one aligned with side plate suture and one aligned<br />

approximately near middle <strong>of</strong> the ambulacral margin<br />

<strong>of</strong> each side plate), with secondary side plate normally<br />

disposed. There are four cover-plate sockets to each side<br />

plate along the main food groove and two or three side<br />

cover-plate sockets along each side <strong>of</strong> a side food groove.<br />

The interradial sutures are depressed and the surfaces <strong>of</strong><br />

the calyx plates are ornamented with finely granular<br />

growth lines parallel to plate margins.<br />

Remarks.—The original types were probably destroyed<br />

in the fire at Indiana <strong>University</strong> in 1883 and<br />

therefore neotypes are here selected.<br />

Occurrence.—Lower Mississippian, upper part <strong>of</strong> Burlington<br />

Limestone, Burlington, Iowa.<br />

Types.—Neotypes, S4,956 (new holotype), S4,955<br />

(new paratype), Springer collection, old no. S3,759, part<br />

<strong>of</strong> 31 specimens, U. S. National Museum, Washington.<br />

EXPLANATION OF PLATE 22<br />

FIGURE<br />

PAGE<br />

1-3. Cordyloblastus roemeri (ETHERIDGE & CARPENTER), plesiotype,<br />

S3,615, Springer coll., U.S. Natl. Mus.; Lower Devonian,<br />

Calizas de Arnao, Le6n, Spain; oral, "D" ambulacral,<br />

aboral views (all X8.2).<br />

4-9. Cordy/oHastus /usitanicus (ETHERIDGE & CARPENTER).<br />

4 6. Plesiotype, S3,6I6, Springer coll., U.S. Natl.<br />

Mus.; Middle Devonian, Gerolstein, Eifel, Germany; oral,<br />

"D" ambulacral, aboral views (all X9.8). 7 9.<br />

Plesiotype, 19,091, Gurley coll., Univ. Chicago; Middle<br />

Devonian, Gerolstein, Germany; oral, aboral, "D"<br />

ambulacral views (all X 11.5) 57

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