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Primary Documents Relating to Darwin and Darwinism<br />

me; and this was long after I had come to Down. The solution,<br />

as I believe, is that the modified offspring of all dominant and<br />

increasing forms tend to become adapted to many and highly<br />

diversified places in the economy of nature.<br />

Source: Francis Darwin, ed., The Life and Letters of Charles Darwin,<br />

Including an Autobiographical Chapter (London: John Murray, 1887),<br />

I: 83–84.<br />

131<br />

Document 6: Darwin to Gray, September 1857<br />

On 5 September 1857, Darwin wrote Asa Gray a letter about<br />

the origin of species. Gray had written a letter to Darwin the previous<br />

month in which Gray stated that his research on plants caused<br />

him to have doubts about the fixity of species. Darwin took the opportunity<br />

presented by Gray’s revelation to describe his theory that<br />

could explain why certain species seemed to be changing. Appended<br />

to the letter was a summary of Darwin’s theory of speciation—the<br />

idea that one species could become the progenitor of two or more<br />

new species. The essay Darwin sent to Gray was as good a summary<br />

of The Origin of Species as Darwin wrote before the book was<br />

published.<br />

After Darwin received Alfred Wallace’s essay entitled ‘‘On the<br />

Tendency of Varieties to Depart Indefinitely from the Original Type’’<br />

in June 1858, he was uncertain about what he should do. Charles<br />

Lyell and Joseph Hooker suggested including the letter and summary<br />

that Darwin sent to Gray along with essays by Wallace and Darwin<br />

about speciation to help prove that Darwin had been working on the<br />

theory before Wallace.<br />

The following extract is a copy of the essay that Darwin sent to<br />

Gray. Darwin corrected the version of essay he had written, and Lyell<br />

and Hooker sent that copy to the secretary of the Linnean Society,<br />

John Joseph Bennett (1801–1876). The major difference between the<br />

two essays was the postscript that Darwin took out of the copy sent<br />

to the Linnean Society.<br />

The essay itself is a fascinating preview of The Origin of Species.<br />

The material that would form important components of Darwin’s<br />

theory—artificial selection by breeders, the importance of geological<br />

time, the cumulative work of natural selection, the struggle for life, geographical<br />

distribution of species, and the importance of classification—<br />

is clearly evident. The examples of the mistletoe and the elephant that<br />

appeared in Chapter III, ‘‘Struggle for Existence,’’ are in the essay. The<br />

problem Darwin had with his use of the term natural selection is also

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