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Book Fauna Palaestina 4 Year 2014 By Prof Dr Norman Ali Bassam Khalaf von Jaffa ISBN 978-9950-383-77-7

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the beetle, and are fixed by the time it emerges from its pupa. The<br />

same applies to the colour, except it may take some days for the<br />

colour of the adult beetle to mature and stabilise. Generally, the<br />

mature colour tends to be fuller and darker than the colour of<br />

the callow (Wikipedia).<br />

Biology<br />

Coccinellids are best known as predators of Sternorrhyncha such<br />

as aphids and scale insects, but the range of prey species that<br />

various Coccinellidae may attack is much wider. A genus of small<br />

black ladybirds, Stethorus, presents one example of predation on<br />

non-Sternorrhyncha; they specialise in mites as prey, notably<br />

Tetranychus spider mites. Stethorus species accordingly are<br />

important in certain examples of biological control (Wikipedia).<br />

Various larger species of Coccinellidae attack caterpillars and<br />

other beetle larvae. Several genera feed on various insects or their<br />

eggs; for example, Coleomegilla species are significant predators of<br />

the eggs and larvae of moths such as species of Spodoptera and<br />

the Plutellidae. Larvae and eggs of ladybirds, either their own or<br />

of other species, can also be important food resources when<br />

alternative prey are scarce. As a family, the Coccinellidae used to<br />

be regarded as purely carnivorous, but they are now known to be<br />

far more omnivorous than previously thought, both as a family<br />

and in individual species; examination of gut contents of<br />

apparently specialist predators commonly yield residues of pollen<br />

and other plant materials. Besides the prey they favour, most<br />

predatory coccinellids include other items in their diets, including<br />

honeydew, pollen, plant sap, nectar, and various fungi. The<br />

significance of such nonprey items in their diets is still under<br />

investigation and discussion (Wikipedia).<br />

Apart from the generalist aphid and scale predators and<br />

incidental substances of botanical origin, many Coccinellidae do<br />

favour or even specialise in certain prey types. This makes some<br />

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