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9 8<br />

MORPHOLOGY<br />

III. RADIATE THALLUS<br />

i. CHARACTERS OF RADIATE THALLUS<br />

In the stratose dorsiventral thallus, there is a widely extended growing<br />

area situated round the free margins of the thallus. In the radiate thallus<br />

of the fruticose or filamentous lichens, growth is confined to an apical region.<br />

Attachment to the substratum is at one point only the base of the plant<br />

thus securing the exposure of all sides equally to light. The cortex<br />

surrounds the fronds, and the gonidia (mostly Protococcaceae) lie in a zone<br />

or in groups between the cortex and the medulla. It is the highest type of<br />

since it secures the widest<br />

vegetative development in the lichen kingdom,<br />

room for the gonidial layer, and the largest opportunity for photosynthesis.<br />

Shrubby upright lichens consist mostly of strap-shaped fronds, either<br />

simple or branched, which may be broadened to thin bands (Fig. 57) or<br />

may be narrowed and thickened till they are almost cylindrical. The fronds<br />

vary in length according to the species from a few millimetres upwards:<br />

Fig. 57. Roccellafuciformis DC.

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