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biologia - Studia

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PLATFORM INITIATION, CLUTCH INITIATION AND CLUTCH SIZE IN COLONIAL GREAT CRESTED GREBES<br />

Clutch initiation dates of all small clutches fell twice into the first decade<br />

of April, 5 times into the second and 8 times into the third. Thus, the number of<br />

small clutches goes seasonally increasing (Table 6).<br />

Platform<br />

Initiation<br />

(D=decade)<br />

Table 6.<br />

Summary of characteristics associated with small clutches in the Reeds<br />

and in the Bulrush.<br />

Clutch initiation with respect to average date for the colony<br />

Earlier On the average date Later<br />

3 4 8<br />

Waiting time from<br />

platform to clutch<br />

initiation<br />

Below Above<br />

average average<br />

Waiting time from<br />

platform to clutch<br />

initiation<br />

Below Above<br />

average average<br />

Waiting time from<br />

platform to clutch<br />

initiation<br />

Below Above<br />

average average<br />

March 1 2 3<br />

1D April 1 1 1<br />

2D April 1 1 1 2<br />

3D April 1<br />

Total 1 2 1 3 1 7<br />

Exceptionally Large Clutches<br />

All 17 full clutches with more than 5 eggs were laid in the Reed zones and<br />

none of them was recorded in 2003 (Table 5). 16 of these clutches were started<br />

prior to the average laying date for the Reeds, 13 of them more than 5 days earlier,<br />

the other 3 just 1 or 2 days earlier. The 17 th clutch was initiated on the average<br />

date. 10 of the really early large clutches knew a platform initiation in March and<br />

they waited for 4, 6, 11, 12, 12, 17, 18, 20, 20 and 20 days before receiving a first<br />

egg. 2 platforms appeared during the first decade of April and waited for 6 days<br />

each before initiating a clutch and one platform was started during the second<br />

decade of April to hold an egg 3 days later. For the 3 clutches initiated just prior to<br />

the average date, one platform appeared in March and received an egg after 26<br />

days, the remaining 2 were initiated in the first decade of April and held an egg 5<br />

and 10 days later. The only clutch started on the average day knew its platform<br />

initiation during the second decade of April and a clutch initiation 4 days later.<br />

15 of the 17 clutches (88%) occurred in nests clumped with at least 3 other<br />

nests and 10 (59%) in nests clumped with at least 6 additional nests. One nest<br />

holding a full clutch of 6 eggs was located in a narrow part of Reed I. It was<br />

clumped with 2 other nests and had its nearest neighbour at 2.3 m. The platform<br />

started on 25 March 2005 received its first egg on 5 April. The last platform was<br />

located at the beginning of Reed II where the stands of vegetation were much<br />

spaced and therefore prohibited having very close neighbours. It was started on 20<br />

53

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