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PETER K. L. NG & DARREN C. J. YEO (2007)<br />

some 1,500 km 2 . However, while the area it is known from appears to be relatively extensive,<br />

it must be noted that the distribution consists of many pockets of populations, and this species<br />

is known to show the greatest variation among the Johora species, facts that point to it probably<br />

being a species complex (Ng 1988). Another probable species complex is the troglophilic<br />

crab, Stygothelphusa bidiensis, which has an unlikely distribution of two disjunct cave systems<br />

in Sarawak (Bau and Gua Serian). The available evidence suggests that the populations in the<br />

two cave systems actually belong to two separate species (unpublished data). The same situation<br />

is true of Lepidothelphusa cognetti, which occurs in the sandstone streams of Bau and Penrissen.<br />

Another point to consider is that for freshwater crabs in developing countries, the line separating<br />

a vulnerable or endangered species is a very fine one. This is mainly because of the very<br />

restricted distributions of many species and the speed of development projects; the time lapse<br />

between project conception and implementation, even for large scale ones, can be as short as<br />

a year.<br />

Using Peninsular Malaysia and Singapore as an example, 42 species of potamids and<br />

parathelphusids are known at present. All the potamids (27 taxa) are found only in Peninsular<br />

Malaysia and Singapore. Of the 15 parathelphusids, 10 are endemic to Peninsular Malaysia<br />

and Singapore, the other five species also occurring in Sumatra or southern Thailand. The<br />

endemic taxa are almost always highland species, or occur on isolated islands. The conservation<br />

of this remarkable diversity is imperative (see also Ng 1988). There is thus, more than ever, a<br />

need to establish more nature reserves and national parks. And careful planning, co-ordination<br />

and supervision to minimise its destructive effects must temper development, inevitable though<br />

it may be. At the same time, other broader, long term issues, those of water-shed conservation,<br />

sufficient size of protected areas, and forest conditions (primary or secondary or disturbed)<br />

must be given due consideration. Such matters if dealt with properly would not just be for the<br />

benefit of freshwater crab diversity, but for the overall ecosystem as well.<br />

ACKNOWLEDGEMENTS<br />

The authors thank the organisers of the workshop for inviting this paper from them, in particular,<br />

Saw Leng Guan, the chair of the organising committee.<br />

REFERENCES<br />

BAHIR, M.M., NG, P.K.L., CRANDALL, K. & PETHIYAGODA, R. 2005. A conservation<br />

assessment of the freshwater crabs of Sri Lanka. Raffles Bulletin of Zoology Supplement<br />

12: 121–126.<br />

CRANBROOK, EARL OF, & FURTADO, J.L. 1988. Freshwaters–Decapod Crustacea. Pp.<br />

225–250 in Earl of Cranbrook (ed.) Key Environments: Malaysia. Pergamon Press, Oxford.<br />

IUCN (International Union for the Conservation of Nature and Natural Resources), 2001. The<br />

IUCN Red List of Threatened Species: 2001 Categories & Criteria. Version 3.1. http://<br />

www.redlist.org/info/categories_criteria2001.html [accessed 01.01.2004].<br />

NG, P.K.L. 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department<br />

of Zoology, National University of Singapore. Shinglee Press, Singapore.<br />

119

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