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CHEY VUN KHEN (2007)<br />

RESEARCH REVIEW<br />

Apart from the taxonomic work mentioned earlier, most of the research work on diversity of<br />

moths and butterflies in Malaysia has focused on their use as bioindicators of habitat quality<br />

and habitat change.<br />

The specialist of Bornean macromoths, Dr. Jeremy Holloway in London, started his association<br />

with Borneo back in 1965 when a Cambridge expedition to Mount Kinabalu was organised. A<br />

paper giving a numerical analysis of the Kinabalu moth samples was published (Holloway<br />

1970), as well as a taxonomic monograph on the moths of Mount Kinabalu (Holloway 1976).<br />

Thereafter, he has been consistently publishing taxonomic monographs in his “Moths of<br />

Borneo” series. Apart from that, he also publishes papers on moth ecology, particularly on the<br />

use of moths as indicators in comparing forest habitats in Malaysia. His papers include one on<br />

the larger moths of Gunung Mulu in Sarawak (Holloway 1984), and the response of moths to<br />

forest conversion in Sabah (Holloway et al. 1992).<br />

Dr. Holloway also trained up two Malaysian entomologists working on moths. One is the<br />

present author who used moths to compare the biodiversity between plantation and natural<br />

forests in Sabah (Chey 1994; Chey et al. 1997), and who later studied the moth diversity of<br />

Lanjak-Entimau in Sarawak (Chey 2000a; 2000b). Another is Dr. Jurie Intachat of FRIM,<br />

who assessed moth diversity in natural and managed forests in Peninsular Malaysia (Intachat<br />

1995; Intachat et al. 1999a; 1999b; 2005). Chey worked on the whole spectrum of macromoths<br />

to inventory biodiversity, while Intachat focused on the geometroid moths specifically to<br />

monitor change.<br />

Others who have worked on moth diversity in Borneo include the German researchers Schulze<br />

& Fiedler (1996, 1997) and Beck et al. (2002).<br />

Research on butterflies as indicators of forest disturbance in Sabah has been carried out since<br />

the mid 1990s by Dr. Jane Hill of the UK and her colleagues. Their papers include species<br />

abundance models (Hill & Hamer 1998), comparison of butterflies in rain forest gaps and<br />

closed-canopy forests (Hill et al. 2001) as well as in natural and selectively logged forests<br />

(Hamer et al. 2003), and the effects of rainfall as opposed to logging on the abundance of a<br />

selected butterfly species (Hill et al. 2003). They used fruit-baited traps, which were also used<br />

to study vertical stratification of activity (Tangah et al. 2004), as well as walk-and-count<br />

ground-based surveys.<br />

In addition to these studies, a lot of general information on moths and butterflies in Malaysia<br />

is provided in the handbook by Holloway et al. (2001) and in the hostplant book by Robinson<br />

et al. (2001).<br />

ACKNOWLEDGEMENT<br />

Dr. J.D. Holloway of the Natural History Museum in London kindly commented on the<br />

manuscript.<br />

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