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CHEY VUN KHEN (2007)<br />
RESEARCH REVIEW<br />
Apart from the taxonomic work mentioned earlier, most of the research work on diversity of<br />
moths and butterflies in Malaysia has focused on their use as bioindicators of habitat quality<br />
and habitat change.<br />
The specialist of Bornean macromoths, Dr. Jeremy Holloway in London, started his association<br />
with Borneo back in 1965 when a Cambridge expedition to Mount Kinabalu was organised. A<br />
paper giving a numerical analysis of the Kinabalu moth samples was published (Holloway<br />
1970), as well as a taxonomic monograph on the moths of Mount Kinabalu (Holloway 1976).<br />
Thereafter, he has been consistently publishing taxonomic monographs in his “Moths of<br />
Borneo” series. Apart from that, he also publishes papers on moth ecology, particularly on the<br />
use of moths as indicators in comparing forest habitats in Malaysia. His papers include one on<br />
the larger moths of Gunung Mulu in Sarawak (Holloway 1984), and the response of moths to<br />
forest conversion in Sabah (Holloway et al. 1992).<br />
Dr. Holloway also trained up two Malaysian entomologists working on moths. One is the<br />
present author who used moths to compare the biodiversity between plantation and natural<br />
forests in Sabah (Chey 1994; Chey et al. 1997), and who later studied the moth diversity of<br />
Lanjak-Entimau in Sarawak (Chey 2000a; 2000b). Another is Dr. Jurie Intachat of FRIM,<br />
who assessed moth diversity in natural and managed forests in Peninsular Malaysia (Intachat<br />
1995; Intachat et al. 1999a; 1999b; 2005). Chey worked on the whole spectrum of macromoths<br />
to inventory biodiversity, while Intachat focused on the geometroid moths specifically to<br />
monitor change.<br />
Others who have worked on moth diversity in Borneo include the German researchers Schulze<br />
& Fiedler (1996, 1997) and Beck et al. (2002).<br />
Research on butterflies as indicators of forest disturbance in Sabah has been carried out since<br />
the mid 1990s by Dr. Jane Hill of the UK and her colleagues. Their papers include species<br />
abundance models (Hill & Hamer 1998), comparison of butterflies in rain forest gaps and<br />
closed-canopy forests (Hill et al. 2001) as well as in natural and selectively logged forests<br />
(Hamer et al. 2003), and the effects of rainfall as opposed to logging on the abundance of a<br />
selected butterfly species (Hill et al. 2003). They used fruit-baited traps, which were also used<br />
to study vertical stratification of activity (Tangah et al. 2004), as well as walk-and-count<br />
ground-based surveys.<br />
In addition to these studies, a lot of general information on moths and butterflies in Malaysia<br />
is provided in the handbook by Holloway et al. (2001) and in the hostplant book by Robinson<br />
et al. (2001).<br />
ACKNOWLEDGEMENT<br />
Dr. J.D. Holloway of the Natural History Museum in London kindly commented on the<br />
manuscript.<br />
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