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THE STATUS OF RESEARCH ON HYMENOPTERA IN MALAYSIA<br />

(approximately 12 mm body length or larger) tended to be higher in disturbed areas. This<br />

could be an example of competitive exclusion, in which certain species are prevented from<br />

occupying an area by the presence of other species (Cox & Moore, 1993). Xanthopimpla<br />

species, as examples of pimpline wasps, lay eggs in suitable hosts, and larger species select<br />

hosts that enable the development of a larger wasp. (Gauld, 1984). Disturbed habitats may<br />

favor the existence of suitable hosts for large Xanthopimpla species, which compete with<br />

smaller species for the limited resources available. Certain large species like X. gampsura<br />

were abundant in disturbed habitats, while X. nigritarsis nigritarsis was found only in pristine<br />

habitats. Although the finding is preliminary and needs to be verified by replication at other<br />

locations, these two species of Xanthopimpla may have the potential to be used as biological<br />

indicators for habitat disturbance.<br />

Table 5. Contingency table 1 for the body length of Xanthopimpla species collected in<br />

undisturbed and disturbed habitats.<br />

Body length (mm) Undisturbed Disturbed<br />

Small (5.30 - 8.66) 16 20<br />

Medium (8.67 - 12.03) 30 11<br />

Large (12.04 - 15.4) 2 11<br />

Total 48 42<br />

1<br />

Chi Square (2 degrees of freedom) = 15.15, p < 0.001.<br />

PUBLICATIONS AND OTHER SOURCES OF INFORMATION ON<br />

ICHNEUMONIDAE AND OTHER PARASITIC HYMENOPTERA<br />

There are many ways to access information on Hymenoptera, in particular the Ichneumonidae<br />

and other parasitic hymenopterans. Provided below is a list of relevant references, revisions,<br />

catalogues, CD-ROMs and electronic information-sources (websites, etc). Useful books for<br />

beginners are those written by LaSalle & Gauld (1993), Gauld & Bolton (1996), Austin &<br />

Dowton (2000), Quicke (1987), Morley (1913) and Goulet & Huber (1993). Books or<br />

catalogues specifically on Braconidae (braconid wasps) and Formicidae (ants) have been written<br />

by van Achterberg (1996), Shenefelt (1975), van der Vecht & Shenefelt (1969) and Bolton<br />

(1997). Many other books or catalogues provide information on Ichneumonid wasps<br />

(Hymenoptera: Ichneumonidae) but these are too numerous to list here. To date, few revisions<br />

on Ichneumonidae and Braconidae have been published; these are Townes (1983), Quicke<br />

(1997) and Simboloti & van Achterberg (1990a & 1990b). Bouèek (1988) and Huang &<br />

Noyes (1994) provide good revisions for Chalcidoidea and Encyrtidae, respectively.<br />

There are several journals that frequently publish or are devoted to research on Hymenoptera<br />

(Table 7). The Oriental Insects Monograph, Pacific Insect Monograph and Ichneumonologia<br />

Orientalis commonly publish articles on Ichneumonidae and Braconidae of the Indo-Australian<br />

and Oriental Regions, while the Zoologische Mededelingen Leiden and Zoologische<br />

Verhandelingen usually publish articles on braconids. The Journal of Hymenoptera Research<br />

publishes research on any aspect of Hymenoptera. Other articles on Hymenoptera diversity<br />

and taxonomy can also sometimes be found in Serangga, Bulletin of Entomological Research,<br />

Biocontrol and various other entomological journals.<br />

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