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CONSERVATION STRATEGIES OF SHOREA LUMUTENSIS (DIPTEROCARPACEAE) IN PENINSULAR MALAYSIA<br />

A total of 416 individuals >1 cm dbh were recorded within the 8-ha plot in Sungai Pinang. The<br />

population density of S. lumutensis >30 cm dbh within the plot was 4.4 trees ha -1 . The prominent<br />

associated species within the habitat are two palm species, i.e., Eugeissona tristis and Calamus<br />

castaneus. The study of the relationship between microtopography and spatial distribution<br />

showed that the species distribution was strongly related to topography; prominent on ridges<br />

and upper slopes, and totally absent in the lower slopes and valleys. This was further supported<br />

by spatial distribution analyses, in which significant spatial aggregation was detected at four<br />

size classes (Fig. 2) and the level of aggregation was highest in SEE and SAP, followed by<br />

POL, and then BIG.<br />

Diameter distribution was skewed, with many more small than large individuals being present.<br />

The distribution was significantly fitted to inverse J-shaped curves (y = ae -bx ; a = 154.6; b =<br />

0.6; r² = 0.98, P < 0.01), indicating abundant regeneration. The medium-sized trees (11–20<br />

cm) constituted 1.7% of the total 416 individuals found within the plot, compared with 8.2% in<br />

the largest-sized trees (>31 cm) and 82.2% in the smallest-sized trees (1–5 cm). Short-term<br />

population dynamics derived from the initial census in September 2001 and a repeat census in<br />

August 2004 showed that a total of 75 trees died over the 3-year study period. Mortality was<br />

detected only at the two lowest-sized classes (1–5 cm and 6–10 cm), 22% and 8% respectively<br />

(Table 1). Growth was slow in most of the trees enumerated, at mean rates around<br />

0.3 mm yr -1 (lowest-sized class) to 2.4 mm yr -1 (highest-sized class) and the mean growth rate<br />

increased with increasing size class.<br />

15<br />

BIG<br />

30<br />

25<br />

POL<br />

10<br />

20<br />

Ripley’s K<br />

5<br />

0<br />

Ripley’s K<br />

15<br />

10<br />

5<br />

0<br />

-5<br />

-5<br />

-10<br />

-10<br />

0 1 2 3 4 5<br />

0 1 2 3 4 5<br />

clas s<br />

Distance class<br />

class<br />

Distance class<br />

70<br />

60<br />

SAP<br />

90<br />

80<br />

SEE<br />

70<br />

50<br />

60<br />

Ripley’s K<br />

40<br />

30<br />

20<br />

10<br />

0<br />

-10<br />

0 1 2 3 4 5<br />

Distance clas class s<br />

Ripley’s K<br />

50<br />

40<br />

30<br />

20<br />

10<br />

0<br />

-10<br />

-20<br />

0 1 2 3 4 5<br />

Distance class class<br />

Fig. 2. Spatial distribution analysis using Ripley’s K-function on four diameter classes of<br />

S. lumutensis within an 8-ha (400 200 m) plot: large trees (BIG >25 cm), pole trees (POL<br />

4–25 cm), saplings (SAP 2.0–2.5 cm) and seedlings (SEE 1.0–1.1 cm). Distance classes were<br />

defined at five intervals, each of 20 m, from 0–20 m (class 1) to 80–100 (class 5). Dotted lines<br />

represent 95% confidence limits.<br />

276

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