(a)swamplaureloakSweet-Florodared horn -maple beam bald shrubsWFigi~re 25. Species composition <strong>of</strong> common trees (>I0 cm dbh; hatched bars) and shrubs andsaplings (2.5-10 dbh; open bars) in two hydric hammocks: (a) Seminole County (data from G. R. Bestand P. Wallace, Center for <strong>Wetlands</strong>, University <strong>of</strong> Florida); (b) northern gulf coastal hammock(Thompson 1980). Importance values (defined in Table 5) were calculated for each size-class withina hammock. Species with importance values <strong>of</strong> 5 or less in both tree and sapling classes wereomitted.Tree saplings dominated the 2.5-10cm dbh :izc class in hydric hammocks<strong>of</strong> Seminole County and the northerngulf coast (Figure 25); shrubs accountedfor less than 20% <strong>of</strong> the basalarea. With the exception <strong>of</strong> cabbagepalm, a1 1 tree species were representedin the smaller-sized category.<strong>The</strong> converse was not true: red maplesapl ings were abundant in the SeminoleCounty hammock, but mature trees <strong>of</strong>that species were absent. In the sameforest, swamp 1 accrel oak, sweetgum,and loblolly pine had higher importancevalues as sapl ings than astrees. Apparently the canopy containedgaps that promoted colonization<strong>of</strong> these shade-intolerant species(Putnam et a7. 1960).Hornbeam and swampbay were more importantas "sapl ings" than as "trees,"because they attain maturity at asmall size (Figure 25). Hornbeam wasan abundant member <strong>of</strong> the subcanopy <strong>of</strong>the Seminole County hammock, the OrangeLakc palm hammock, and the inlandreach <strong>of</strong> Gulf Hammock, and it musthave dominated this stratum in theHi1 1 sborough River hydric hammock(Table 5). However, this species wasscarce in the northern gulf coastalhammock (Thompson 1980) and non-existentin many <strong>of</strong> the hydric hammocksthat we visited, including SilverSprings, Wekiva Springs, Myakka River,and Sanchez Prairie (in San FelascoHammock State Preserve). Hornbeam'sabsence from the last forest is puzzling,since the species is common insurrounding mesic hammock.Yaupon, wax-myrtle, and dahoon werethe most common shrubs on the northerngulf coast (Thompson 1980), whereasswamp dogwood and wax-myrtle were theonly shrubs sampled in the SeminoleCounty hydric hammock (G. R. Best andP. Wall ace, pers. comm. ). Wax-myrtle
was the most frequent shrub in the hydrichanimccks that we surveyed. Fewsnruo species uLLur red r eyui dr ;y, dr~dseveral were restr-~cted to one or afew hammocks. Blue-stem palmetto appearedin several stands within theSi 1 ver Springs hydric hammock. Needlepalm was present only at WekivaSprings and Tiger Creek. <strong>The</strong> drierparts <strong>of</strong> the Sanchez Prairie hydrichammock featured blueberry bushes(Vaccinium spp. ) , but the wetter areashad no shrubs. Among the hydric hamiiid~khwe ;l is i Led, biil u?. veyetatf \u':~densest at Wekiva Springs.tiydric hammocks with a high abundance<strong>of</strong> cabbage palm, such as Tosohatcheeand Myakka River (Table 5),generally had few plants in the shrublayer, except for very young palms,and no ground vegetation (Figure 18).<strong>The</strong> extent <strong>of</strong> the ground layer was extremelyvariable both among and withinthe reinainiiig fcr-ests. In the SilverSprings hydric hammock, dense patches<strong>of</strong> spi kegrasses (Chasmanthium spp.)inhabited clearings. Vegetation,mainly grasses and ferns, covered from74% to 96% <strong>of</strong> the ground in six standsat the northern end <strong>of</strong> the Gulf Hammockregion (Thompson 1980). <strong>The</strong>pl ants were present in sl ightl y openareas and absent from low, wet places,suggesting that they were 1 imi ted bylow 1 ight and extended f1 ooding.Herbaceous vegetation was most lushand diverse in the Tiger Creek, SilverSprings, and in1 and Gulf Hammockforests. In those hydric hammocks, apr<strong>of</strong>usion <strong>of</strong> ferns, grasses, sedges,and herbs covered the ground (Figure19).<strong>The</strong> largest contiguous stand <strong>of</strong> hydrichammock, known locally as GulfHammock, comprises the section <strong>of</strong> gulfcoastal hammock between the Suwanneeand With1 acoochee Rivers. Originallycovering more than 40,000 ha, GulfHammock has been reduced by clearingfor pine plantations and agriculture(Simons et dl. 1988). Gulf Hammock ishighly diverse, encompassing most <strong>of</strong>the variation in structure and composi tion found among hydric hammocks.Few hammocks have such a complete setaf the more common plant species, nor<strong>of</strong> the rare ones, such as cedar elm,American plum, pink-root, pine-wooddaintiss, hiid the two indian plantainsCaca7 i a suaveolens and Arnog7ossum diversifolium.<strong>The</strong> interface <strong>of</strong> salt marsh and GulfHammock forest (Figure 26) is very irregular.Many islands <strong>of</strong> hammock arefound in the marsh, and the marsh extendsinto the forest along tidalcreeks, sometimes for considerabledistances. In most places along theboundary, the forest begins abruptlywith a dense, 12- 15-meter-tall stand<strong>of</strong> cabbage palm, live oak, southernred-cedar, and occasionally water locust.Loblolly pine sometimes occursat the forest edge, and in a few areas<strong>of</strong> coastal Gulf Hammock, it dominatesthe forest, <strong>of</strong>ten in association witha dense ground cover <strong>of</strong> St. AugustineFigure 26. Edge <strong>of</strong> hydric hammock and saltmarsh in Gulf Hammock, along the Gulf <strong>of</strong> Mexiconear the Withlacoochee River, Citrus County,Florida.
- Page 2 and 3: Copies of this publication may be o
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Wharton, C.H. 1977. The naturalenvi