Upper Myakka Lake/-HAMMOCKINVASION0.5 kmFigure 31. Expansion <strong>of</strong> hydric hammock(Shep's Island) into freshwater marsh, MyakkaRiver State Park, 1957-1972.hammock is composed <strong>of</strong> cabbage palmand live oak, the invading vegetationconsists <strong>of</strong> swamp laurel oak, Floridaelm, and, closer to the marsh, waterlocust and pop ash (Figure 32). Severeflooding during the summer <strong>of</strong>1982 killed many swamp laurel oaks,but not 1 ive oaks. Apparently thelatter species can withstand occasionaldeep inundation but not prolongedsoi 1 saturation.3.3.3 Eda~hic ConditionsParticular plant species may be favoredin some hydric hammocks because<strong>of</strong> variations in soil type. Changesin soil texture, organic content, andpH can markedly affect drainage andnutrient avail abil i ty, and therefore,the growth and survival <strong>of</strong> variousspecies. For example, sweetbay generallyis associated with organic soils(Wharton et a7. 1976). This speciesis most abundant in hydric hammockssuch as the Wekiva Springs forest(Table 5) that are characterized byseepage <strong>of</strong> ground water and organicrichsoils. <strong>The</strong> abundance <strong>of</strong> lob101 lypine in the northern gulf coastal hammocksi s negatively correl ated wi thsoil pH (Thompson 1980). Among thehydric hammocks sampled in Table 5,loblolly pine was most important inthe Silver Springs forest; the clayeysoils in that region are more acidthan the fine sands common to most hydrichammocks. <strong>The</strong> abundance patterns<strong>of</strong> water oak, both within the northernGulf Hammock and among hydric hammocks,parallel those <strong>of</strong> loblollypine, suggesting that growth <strong>of</strong> bothspecies is favored by similar conditions.CABBAGE PALMLIVE OAKSWAMP LAURELOAKFLORIDA ELMWATER LOCUSTPOP ASHFigure 32. Pr<strong>of</strong>ile <strong>of</strong> expanding hydric hammock, Myakka River State Park.
<strong>The</strong> salt concentration <strong>of</strong> hydrichammocksoi 1 s strongly influences communI ty SxruLLur e. i~drill!-mediately adjacent to salt marshes <strong>of</strong>the central gulf coast are composedalmost entirely <strong>of</strong> Cabbage palm,southern red-cedar, and live oak(Figure 21).termed coastal<strong>The</strong>se forests werehammocks by Swindell(1949), the Florida Game and Fresh WaterFish Commission (1976), andClewell (1981), but we consider themto be an extreme type <strong>of</strong> hydric hammockLi,ai, ;> > iiiipl i fie6 Ly o~ed;i >prayand floodwater. <strong>The</strong> dominance <strong>of</strong> cabbagepalm in association with 1 ive oakand red-cedar continues for about 2 kminland (Figure 33). As distance fromthe sal tmarsh boundary increases, theforest is enriched with additionalspecies. Loblolly pine is the firstaddition, sometimes occurring at theforest edge. Cedar elm occurs asscattered trees beginning perhaps 200-300 in froia the coast. At dbout 1 kn~,Florida elm, sugarberry, and Floridamaple become common in some places.Sweetgum, swamp laurel oak, and hornbeamare abundant at 2 km. Averagei~ ~I~I,~I~IUL~>maximum height <strong>of</strong> trees increases from12-14 m at the marsh edge to 18-20 mitridnri i.5 km. f hompson (193'3) notedthat the structure <strong>of</strong> hydric-hammockstands in the northern Gulf Hammockalso varied with distance from tidalmarsh. Cabbage palm and southern redcedardecreased i n re1 ative importanceand sweetgum increased, as stands werelocated farther from the marsh andsoil concentrati ons <strong>of</strong> solubl e saltsdecreased. Cabbage palm, 1 ive oak,and southern red-cedar are very salttoleititit{Kurr and Wagner 1957). <strong>The</strong>last species has been termed a calciphile(calcium-1 oving) because it a1 -most always is associated with 1 imestoneoutcrops or coastal (salty) regions.However, southern red-cedardoes not require salt; it may be restrictedto sa1 t-influenced hydrichammocks because it is excluded fromothers by less sal t-tolerant species.Absent from many hydric hammocks,southern red-cedar is most abundant atthe seaward edge <strong>of</strong> the gulf coastalhammocks and in hydric hammocks <strong>of</strong> theupper St. Johns River basin (Table 5).<strong>The</strong> species composition <strong>of</strong> hydric hammocksin these two areas is very similar:almost exclusively cabbage palm,live oak, and southern red-cedar. <strong>The</strong>hydric-hammock soi 1 s adjacent to themarshes <strong>of</strong> the St. Johns River arebathed by occasional river overflowcontaining salts originating as upstreamartesian flow <strong>of</strong> fossil sea water(see Chapter 2.3).3h0 6i0 960 12b0 15b0 1800 2 100 2400DISTANCE FROM SALT MARSH (m)Figure 33. Change in tree-species composition<strong>of</strong> a hydric hammock with increasing distancefrom its salt-marsh boundary. <strong>The</strong> point-quartermethod was used, at 50-rn intervals, to Sample thetrees along a transect in Gulf Hammock.<strong>Hammocks</strong> commonly are thought to befree <strong>of</strong> fire and, indeed, to owe theirexistence to protection from this disturbance(e.g. , We1 1 s 1942). Harper(1915) noted <strong>of</strong> hydric hammocks: "<strong>The</strong>humus probably seldom or never getsdry enough to burn, so that fire doesnot need to be reckoned with." He emphasizedtwo features <strong>of</strong> this communitythat tend to reduce fire frequencyand intensity: wet soils and(usually) the lack <strong>of</strong> a substantial1 i tter 1 ayer. However, we observedfire scars in virtually every hydrichammock we visited. Almost all <strong>of</strong> thecharred trunks were <strong>of</strong> cabbage palms.Fire mark5 are preserved on these
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