locally collected litter lost 85% <strong>of</strong>its mass in one year at a streambanksite, but only 58% annually in theterrace forest dominated by oaks(Shure et a]. 1985). <strong>The</strong> leaf decompositionrates <strong>of</strong> most floodplainspecies, including tupelo, ash, blackgum, red maple, and sweetgum, werevery rapid, but those <strong>of</strong> bald cypressand swamp laurel oak were considerablyslower (Elder and Cairns 1982; Shureet a?. 1985). Leaves that were recalcitrant to decomposition tended tohave higher carbon:nitrogen ratios andgreater concentrations <strong>of</strong> 1 ignin andcell ulose. Decomposition rates <strong>of</strong> oak1 eaves besides swamp 1 aurel were notmeasured, but they also are 1 ikely tobe slow because <strong>of</strong> similar chemicalcharacter] st~cs. Ai rnougn i i r;r;er decompositionis likely to be slow inhydric hammocks because <strong>of</strong> pl antspecies composition, low flooding frequency,and the absence <strong>of</strong> strong waterflow, probably only a small amount<strong>of</strong> 1 itter and decomposed material iswashed out by occasional floods. <strong>The</strong>amount and form (part icul ate, dissolved)<strong>of</strong> export from a hammock dependson current velocity, the timing<strong>of</strong> floods in reiation to the seasonalpattern <strong>of</strong> litter fall, and uptakerates <strong>of</strong> dissolved nutrients by hammockplants.
CHAPTER 4. ANIMALS4.1 INTRODUCTIONVirtually nothing has been publishedabout invertebrates inhabiting hydrichammock. This habitat is important tocertain butterfl ies (John A. Fluno,Department <strong>of</strong> Entomology, Ohio StateUniversity, retired; pers. comm.) ,Sugarberry is the sole host plant forthe snout butterfly (Libytheana bachmanii), the hackberry butterfly(Asterocampa celtis), and the tawnyemperor (Asterocampa clyton) . Sugarberryand water elm (P7anera aguatica)are primary hosts for the questionmarkbutterfly (Polygonia interrogat ionis),with false nettle as an occasionalhost. Caterpillars <strong>of</strong> the easterntiger swallowtail (Papilio glaucus)feed on Fraxinus, L iriodendron, andNagno7 i a virginiana.Very few animals are endemic to hydrichammock. <strong>The</strong> crayfish Procambarusgeodytes, a primary burrower,appears to have an endemic distributionin the hydric hammock along SilverRiver and in other forested wetlands<strong>of</strong> the Oklawaha River watershed(Franz 1976). A subspecies <strong>of</strong> thesoutheastern shrew (Sorex longirostriseionis) was describ.ed by Davis (1957)from the hydric hammock around HomosassaSprings, Citrus County,Florida. However, the actual distri -bution <strong>of</strong> this form and its validityas a taxon are uncertain (Humphrey etal. 1986). Despite their commonnames, neither the Gulf Hammock ratsnake nor the Gulf Hammock dwarf sirenare restricted to hydric hammock,though the rat snake occurs there; thesiren occurs in ponds in and beyondhydric hammock.Though the biology <strong>of</strong> most vertebratesfound in hydric hammock isfairly well known, very few have beenstudied there. <strong>The</strong> vertebrate fauna<strong>of</strong> hydric hammock is not unique to thehabitat, resembl ing faunas <strong>of</strong> mostother forested habitats in peninsularFlorida. However, the hydric-hammockfauna is luxuriant. Compared withother types <strong>of</strong> forest in the region,hydric hammock has a highly diversevertebrate fauna and a high abundance<strong>of</strong> selected species.4.2 REPTILES AND AMPHIBIANS<strong>The</strong> herpet<strong>of</strong>auna has no speciesunique to hydric hammock and is representative<strong>of</strong> the region (Table 7).Characteristic species include southernblack racer, rat snake, Floridabox turtle, green anole, ground skink,broad-headed skink, southern toad,green treefrog, squirrel treefrog, andeastern narrow-mouth toad. <strong>The</strong>coastal variant <strong>of</strong> hydric hammocklacks many species found in the inlandvariants <strong>of</strong> hydric hammock wherefloodwater is fresh rather than brackish.<strong>The</strong> blue-striped ribbon snakeand blue-striped garter snake arewidespread in several habitats <strong>of</strong> thegulf coastal region, but they werefound only in the coastal sample <strong>of</strong>hydric hammock shown in Table 7. Inland hydric hammock has an association<strong>of</strong> reptiles and amphibians that appearsto be ecotonal between mesichammock and swamp forest. Lob1 01 lypine-dominated hydric hammock includesa number <strong>of</strong> species characteristic <strong>of</strong>pine flatwoods (pinewoods snake, scarletking snake, pinewoods treefrog).Some quantitative data are avail ableon the reptile and amphibian community
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DISCLAIMERThe opinions and recommen
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CONVERSION TABLEMetric to U.S. Cust
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FIGURESNumber1AL...............Dist
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NumberTABLESPaqeClassifications of
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