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The Ecology of Hydric Hammocks - USGS National Wetlands ...

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grass. Sal t-tolerant shrubs such as1r.a frotescens (marsh elder), Lycrrtmcarol inianum (Chri stmasberry), andBaccharis angust if07 ia, B.g70merulifolia, and B. ha7imifolia(saltbushes) are abundant at the forestedge in some locations. Vines arerare at the edge, and both vines andshrubs are scarce in the forest interioruntil at least 1 km inland. <strong>The</strong>ground cover is very sparse. Thiscoastal part <strong>of</strong> Gulf Hammock is quitesimil ar to the hydric hammocks borderingthe marshes along the St. Johnsand Myakka Rivers.With increasing distance from thesalt marsh, cabbage palm, live oak,and southern red-cedar decl ine andhardwoods increase in dominance. Beginningat about 2 km inland, GulfHammock is divisible into three mainvegetation types. Swamps <strong>of</strong> bald cypress,red maple, swamp tupelo, andgreen ash occur in low areas and alongthe poorly defined creek drainages.Forests that might be considered eithermesic or hydric hammock occupysl ightly elevated ridges. <strong>The</strong> overstorycontains swamp chestnut, Shumard,live, laurel, and water oaks,plus sweetgum, southern magnol i a, sugarberry,winged elm, Florida elm,Florida maple, lob101 ly pine, southernred-cedar, pignut hickory, persimmon,red mulberry, and basswood. <strong>The</strong> understoryand ground cover also are diverse.<strong>The</strong> third type <strong>of</strong> vegetation,the majority <strong>of</strong> Gulf Hammock, is betweenthese two "extremes" in speciescomposition and is clearly hydric hammock.This major hydric hammock (Figure19) consists mainly <strong>of</strong> swamp laureloak and sweetgum in combination withlive oak, water oak, loblolly pine,Florida elm, basswood, persimmon, redmaple, sweetbay, sugarberry, and cabbagepalm. <strong>The</strong> average height <strong>of</strong> thedominant canopy trees is about 30 m,with scattered lob1 01 ly pines emerging3-6 m above the canopy. <strong>The</strong> scatteredlive oaks are by far the largesttrees, averaging about twice the trunkdiameter and crown spread <strong>of</strong> the othercanopy tree species. <strong>The</strong> ground atthe bases <strong>of</strong> these big live oaks <strong>of</strong>tenis raised by the root system to form amound. This microtoaoara~hv is particularlywell suited for the estab-1 ishment <strong>of</strong> magnolias, and, in conjunctionwith the spreading crown <strong>of</strong>the live oak, for the growth <strong>of</strong> severalspecies <strong>of</strong> vines (bullace andsummer grape, pepper vine, rattanvine, and climbing buckthorn). A subcanopydominated by hornbeam, wax-myrtle,swamp dogwood, and various treesap1 ings <strong>of</strong>ten is we1 1 developed. <strong>The</strong>ground layer commonly is a dense mixture<strong>of</strong> grasses, sedges, and ferns,but in wet areas the cover is mostlyleaf 1 itter with only a scattering <strong>of</strong>herbaceous pl ants.Patches <strong>of</strong> hydric hammock dominatedby loblolly pine are found within GulfHammock. Some <strong>of</strong> these are naturaland others are the result <strong>of</strong> human activity.Some <strong>of</strong> the higher ridgeswere cleared for farmjng 1 ong ago, apparentlyby German immigrants, andthen abandoned. Now stands <strong>of</strong> largeloblolly pines, locally known as"German Islands", cover the oldfields. A modern activity with similarresults is the clearcutting <strong>of</strong> extensiveareas <strong>of</strong> Gulf Hammock foll owedby the planting <strong>of</strong> loblolly pine.Gulf Hammock and its surroundingsare very flat, with one exception.Old sand dunes .covered with sand pinescrub vegetation are found on thenorth side, just inland from CedarKey. A mixture <strong>of</strong> swamp and hydrichammock occurs adjacent to the dunes,but both <strong>of</strong> these types are somewhatdifferent in composition than elsewherein Gulf Hammock, presumably dueto the continuous supply <strong>of</strong> waterseeping out <strong>of</strong> the dunes and to thethicker layer <strong>of</strong> organic muck. Swamptupelo, the dominant tree or codominantwith green ash, is much moreabundant than in other swamps <strong>of</strong> theregion. <strong>The</strong> hydric hammock also isdifferent in its abundance <strong>of</strong> needlepalm, which is quite scarce elsewherein Gulf Hammock. This part <strong>of</strong> GulfHammock is most similar to the hydrichammocks at Wekiva Springs, MormonBranch Botanical Area in the Ocala <strong>National</strong>Forest, and Tiger Creek.

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