366 PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES Series 4, Volume 63, No. 12 Yunnan Province. The Gaoligong Shan forms the southwestern limit of the known distributional range of the genus. Key for Identification of Subgenera of Queinnectrechus in the Gaoligong Shan Region 1. Pronotum with basal angles prolonged posteriorly as slender digitiform processes (Figs. 12a, 13a) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Subgenus Queinnectrechus Deuve 1’ Pronotum with basal angles simple (Fig. 14a). . . Subgenus Gaoligongtrechus subgen. nov. Subgenus Queinnectrechus Deuve, 1992 Queinnectrechus Deuve, 1992a:354. DIAGNOSIS.— Adults of this subgenus (Figs. 12a, 13a) can be recognized by the following combination of character states: size moderate (BL = 3.5 to 4.8 mm), apterous, body dark, reddish brown to piceous, surface micaceous; pronotum with basal angles prolonged posteriorly as slender digitiform processes, with two setae (midlateral and basolateral) present on each side; elytra markedly convex, inflated, slightly tear-shaped, humeri effaced, elytral discal striae effaced, with two (in most members) or three (in a few members) discal setae present, aligned on interval 3 near stria 3, preapical seta absent from most members. GEOGRAPHICAL DISTRIBUTION.— Same as for genus (see above). Key for Identification of Adults of Subgenus Queinnectrechus species of the Gaoligong Shan Region 1. Size larger (BL = 4.3 to 4.8 mm), elytra (Fig. 12a) more elongate; median lobe of male aedeagus broadest at mid-shaft, with apex long and recurved. . . . . . . . . . . . . Q. griswoldi sp. nov. Size smaller (BL = 3.5 to 3.8 mm), elytra (Fig. 13a) shorter, ovoid; median lobe of male aedeagus slender, with apex short and not at all curved. . . . . . . . . . . . . . Q. gongshanicus sp. nov. Queinnectrechus (s. str.) griswoldi Deuve and Kavanaugh, sp. nov. (Figs. 12, 37b, 38a, 38b, 39b, 46–48) TYPE MATERIAL.— Holotype, a male, in IOZ, labeled: “CASENT 1026334”/ “CHINA, Yunnan, Gongshan County, Cikai Township, 0.1 km SE of Heipu Yakou in valley below tunnel, N27.76978°/ E98.44681°,”/ “3720 m, 13 August 2006, Stop #DHK-2006-073 D.H. Kavanaugh & J.A. Miller collectors”/ “HOLOTYPE Queinnectrechus (s.str.) griswoldi Deuve & Kavanaugh, sp. nov. designated 2016” [red label]. Paratypes (a total of 20): 5 males and 3 females (in CAS, IOZ, MNHN) labeled same as holotype except first label “CASENT 1026333”, “CASENT 1026335” to “CASENT 1026338” and “CASENT 1026330” to “CASENT 1026332”, respectively; 1 female (in IOZ) labeled “CASENT 1010344”/ “CHINA, Yunnan, Gongshan County, Cikai Township, 52.6 km w of Gongshan on Dulong Valley Road, 3360-3380 m,”/ “N27.77032°/ E098.44661°, 1-2 October 2002, Stop #DHK-2002-034, D.H. Kavanaugh & P.E. Marek collectors”; 1 male and 3 females (in CAS, IOZ) labeled “CASENT 1024862” and “CASENT 1024863” to “CASENT 1024865”, respectively/ “CHINA, Yunnan, Gongshan County, Bingzhongluo Township, Sw slope of Kawakarpu Shan, on slope NE of Chukuai Lake, 3950,”/”N27.98206°/ E098.48027°, 20 August 2006, Stop #DHK-2006-086 Y. Liu, P. Hu, D.Z. Dong, & J. wang collectors”; 3 males and 1 female (in CAS, IOZ, MNHN) labeled “CASENT 1026813” to “CASENT 1026815” and “CASENT 1026815”, respectively/ “CHINA, Yunnan, Gongshan County, Bingzhongluo Township, Sw slope of Kawakarpu Shan at Chukuai Lake, 3720 m,”/ “N27.98121°/ E098.47580°, 18 August 2006 Stop #DHK-2006-079 J.A. Miller, D.Z. Dong, & Y. Liu collectors”; 1 female (in CAS) labeled
DEUVE ET AL.: CARABID BEETLE FAUNA OF THE GAOLIGONG MOUNTAINS 367 FIGURE 12. Queinnectrechus (s. str.) griswoldi sp. nov.; a. Dorsal habitus (CASENT1026334). b. Median lobe of aedeagus of male (CASENT1026334), left lateral aspect. c. Map of locality records (red circles) for Q.griswoldi in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.