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310 PrOCeedINgS OF THe CALIFOrNIA ACAdeMy OF SCIeNCeS Series 4, Volume 63, No. 8 Mimulus verbenaceus Greene (M. cardinalis Benth. var. verbenaceus (greene) Kearney & Peebles) Native perennial herb; SSM. DB 4527, Fig. 12. This species has been treated recently as Erythranthe verbenacea (greene) Nesom & Fraga (Barker et al. 2012). Other collections of Mimulus with red flowers have been made in the southern Black Mountains (e.g., Christy 1494, Goodding & Reeder 22, and Walker & Prigge s.n.; all at ASU and seen on SeINet 2016) and show morphological similarities to both M. cardinalis and M. verbenaceus. These two species have been treated as conspecific and the latter has been recognized as a variety of the former. Molecular studies (Beardsley et al. 2003) reveal the species to be distinct and not phylogenetically sister to one another. examination of materials at CAS/dS show that most specimens can be distinguished only by the orientation of the anther thecae: parallel to sagittate (i.e., those of the pair spreading from the filament at an angle ≤ 45° and thus arrowhead shaped) in M. verbenaceus versus sagittate (but with the pair spreading from the filament at an angle ˃ 45°) to explanate (i.e., those of a pair oriented apex to apex and ± perpendicular to filament) in M. cardinalis. Other characters sometimes used to distinguish these species (e.g., corolla tube length from base of sinus between calyx lobes to separation of limb from tube) varies among specimens from the flora area, and do not correlate with anther orientation. The collections from the flora area cited above all have anther thecae as noted for M. verbenaceus. Plantaginaceae Keckiella antirrhinoides (Benth.) Straw subsp. microphylla (A. Gray) N.H. Holmgren Native shrub; HdwS, LdwS, MdS. B 8931; DB 5644, 5990. Maurandella antirrhiniflora (Willd.) Rothm. (Maurandya antirrhiniflora willd.) Native perennial vine; HdwS, MdS. B 8967; DBH 5736. Mohavea confertiflora (Benth.) A. Heller Native winter ephemeral herb; CBS, LdwS. DB 4663, 6404; DBa 5168. Fig. 3. Neogaerrhinum filipes (A. Gray) Rothm. Native winter ephemeral herb (vine-like); CBS, LdwS. DB 4656, 4692; DBa 5145. Penstemon eatonii A. Gray subsp. undosus (M.E. Jones) D.D. Keck Native perennial herb; HdwS, MdS. B 9000, 9358; DB 5923, 6494. Penstemon palmeri A. Gray Native perennial herb; PJw. DB 6705. Although this fruiting collection lacks flowers, the plant’s overall size (to 2 m) is more characteristic of P. palmeri than of P. pseudospectabilis or P. bicolor. Plantago ovata Forssk. Native winter ephemeral herb; CBS, HdwS, LdwS, MdS. DB 4634, 4669; DBa 5121. Plantago major L. Introduced perennial herb; SSM. DB 5998. Veronica anagallis-aquatica L. Introduced perennial herb; SSM. B 9366; DB 6448. Poaceae Achnatherum ×bloomeri (Bol.) Barkworth Native perennial herb; MdS. Michaels 2091 (ArIz). Achnatherum coronatum (Thurb.) Barkworth (Stipa coronata Thurb. var. depauperata (M.e. Jones) Hitchc.) Native perennial herb; HdwS, MdS. DB 4543; DBH 5688. Achnatherum hymenoides (Roem & Schult.) Barkworth (Oryzopsis hymenoides (roem. & Schult.) ricker) Native perennial herb; BBS, MdS, PJw. DB 6477. Andropogon glomeratus (Walter) Britton, Sterns & Poggenb. Native perennial herb; SSM. DB 4555. Aristida adscensionis L. Native year-round ephemeral herb; HdwS, MdS. DB 4549, 6307. Aristida purpurea Nutt. var. parishii (Hitchc.) Allred (A. parishii Hitchc.) Native perenni-
BUTTerwICK & dANIeL: FLOrA OF SOUTHerN BLACK MTS., ArIzONA 311 al herb; HdwS, MdS. B 8999; DB 4600, 5722. Aristida purpurea Nutt. var. nealleyi (Vasey) Allred (A. purpurea var. glauca (Nees) A.H. Holmgren & N.H. Holmgren) Native perennial herb; CBS, HdwS, LdwS, MdS, PJw. B 8979, 9333; DB 4505, 4593, 5602, 5643, 6308. Avena barbata Pott ex Link Introduced winter ephemeral herb; MdS. Reeder & Reeder 8986 (ArIz). Avena fatua L. Introduced winter ephemeral herb; CBS. B 9346; DB 6011. Bothriochloa barbinodis (Lag.) Herter Native perennial herb; HdwS, MdS. B 8944; DB 4575, 4627. Bouteloua aristidoides (Kunth) Griseb. Rominger 319-65 (ASC). Bouteloua barbata Lag. var. barbata 4619, 5983. Native summer ephemeral herb; CBS, MdS. Native winter ephemeral herb; CBS, HdwS. DB Bouteloua curtipendula (Michx.) Torr. var. caespitosa Gould & Kapadia Native perennial herb; HdwS, MdS, PJw. DB 4571, 5974; DBH 5697. Bromus arizonicus (Shear) Stebbins Native winter ephemeral herb; HdwS. B 8903; DBP 5956. Bromus berteroanus Colla (B. trinii desv.) Native winter ephemeral herb; HdwS, MdS. DB 5625, 5648; DBH 5774; Reeder & Reeder 8984 (ArIz, ASU). Bromus carinatus Hook. & Arn. Native perennial herb; HdwS. DBH 5694. Bromus diandrus Roth (B. rigidus roth) Introduced winter ephemeral herb; dOw, MdS. B 9356, 9362. Bromus rubens L. Introduced winter ephemeral herb; common throughout. DB 4610. Bromus tectorum L. Introduced winter ephemeral herb; MdS. Reeder & Reeder 8987 (ArIz). Cynodon dactylon (L.) Pers. Introduced perennial herb; SSM. DB 4529, 6454. Dasyochloa pulchella (Kunth) Willd. ex Rydb. (Erioneuron pulchellum (Kunth) Tateoka) Native perennial herb; BBS, HdwS, MdS. DB 4615. Digitaria californica (Benth.) Henrard (Trichachne californica (Benth.) Chase) Native perennial herb; HdwS, MdS. DB 4573. Echinochloa colona (L.) Link Introduced summer ephemeral herb; SSM. DB 6313. Elymus elymoides (Raf.) Swezey subsp. elymoides (Sitanion hystrix (Nutt.) J.g. Sm.) Native perennial herb; HdwS, MdS. DB 6393, 6484. Elymus elymoides subsp. brevifolius (J.G. Sm.) Barkworth (Sitanion hystrix var. brevifolium (J.g. Sm.) C.L. Hitchc.) Native perennial herb; HdwS, MdS. DB 6310; DBH 5716. Elymus multisetus (J.G. Sm.) Burtt Davy (Sitanion jubatum J.g. Sm.) Native perennial herb; BBS, MdS. Gould & Darrow 4294 (ArIz). Hilaria rigida (Thurb.) Benth. Native perennial herb; BBS, HdwS, LdwS, MdS, PJw. DB 4684; DBH 5691. Hordeum murinum L. subsp. glaucum (Steud.) Tzvelev (H. glaucum Steud.) Introduced winter ephemeral herb; SSM. B 9363; DB 5680, 6493. Hordeum pusillum Nutt. Native winter ephemeral herb; SSM. DBH 5768. Imperata brevifolia Vasey Native perennial herb; SSM. DB 4568. Jarava speciosa (Trin. & Rupr.) Peñail. (Stipa speciosa Trin. & rupr.) Native perennial herb; BBS, HdwS, LdwS, MdS, PJw. B 9350; DB 4524, 4611, 5655; Hevly & Hovezak 58 (deS); Reeder & Reeder 8985 (ArIz).
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