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DEUVE ET AL.: CARABID BEETLE FAUNA OF THE GAOLIGONG MOUNTAINS 369<br />
Elytra. Elytral silhouette slightly tear-shaped, with humeri effaced, disc markedly convex,<br />
smooth and glabrous. All discal striae effaced except for faintly impressed parascutellar striole and<br />
a short, faintly impressed recurrent stria. Basal setiferous pore present. Three discal setae present,<br />
aligned along presumed track of stria 3. Preapicale seta absent, one or two apicoangular setules<br />
present. Umbilicate setal series with setae of humeral group equidistance from each other, with the<br />
first slightly more medially inserted than the others, setae of median group inserted distinctly posterior<br />
to middle.<br />
Legs. Moderately long but slender. Protibiae with longitudinal furrow, sparely pubescent apically<br />
on anterior surface. Male protarsomeres 1 and 2 dilated and apicomedially toothed.<br />
Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and<br />
ventrite VII of males apically with one pair of paramedial setae, females with two pairs.<br />
Male aedeagus. Median lobe (Fig. 12b) large, elongate, broadest at mid-shaft, with apex long<br />
and recurved dorsally; endophallus with internal sclerites acuminate apically.<br />
COMMENTS.— Based on similarities in the form of the aedeagus of males, this species is closely<br />
related to Queinnectrechus jiuhecola Deuve & Kavanaugh (Deuve et al. 2015), described from<br />
Lijiang County, northwestern Yunnan. However, members of this new species are distinguished<br />
from those of the latter by their much smaller size (BL = 5.3 to 5.8 mm in Q.jiuhecola members),<br />
more elongate body form, more convex elytra, discal striae fully effaced (at least three medial striae<br />
evident in Q.jiuhecola members) and, most significantly, preapical seta absent (present near<br />
stria 2 subapically in Q.jiuhecola members).<br />
In a recent paper, Belousov and Kabak (2016) established a new genus, Uenoites, in which<br />
they placed Q.jiuhecola, as well as three other species previously included in Kozlovites or Deuveotrechus<br />
Uéno (1995), based on the presence of a preapical seta, more convex tempora, and the<br />
median lobe of the male aedeagus not markedly hooked apically. However, all these features are<br />
plesiomorphic among trechines, which is problematic for demonstrating phylogenetic affinity of<br />
the included taxa. Clearly, phylogenetic relationships among these groups of species remain unresolved.<br />
As noted above, the genitalia of Q.griswoldi males are very similar to those of Q.jiuhecola<br />
males, wherease those of the new species described below, Queinnectrechus gongshanicus sp.<br />
nov., are of a very different form, although both of the new species described here are members of<br />
genus Queinnectrechus.<br />
HABITAT DISTRIBUTION.— Members of this species have been found in a variety of microhabitats<br />
in the alpine zone near the crest of the Gaoligong Shan and the eastern slope, at elevations<br />
ranging from 3270 to 3950 m. Specimens were collected under stones in moist meadows and on<br />
tundra slopes and ridges with sparse to thick herbaceous vegetation (Figs. 37b, 38a, 38b, 39b), at<br />
the edges of small streams and seeps from talus slopes, and at the edges of Rhododendron thickets<br />
up to 1.5 m tall. Members of this species have been found syntopic with specimens of Queinnectrechus<br />
(Gaoligongtrechus) balli sp. nov., Queinnectrechus (s.str.) gongshanicus sp. nov., Trechus<br />
gongshanensis sp. nov., Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov.<br />
GEOGRAPHICAL DISTRIBUTION WITHIN THE GAOLIGONG SHAN.— Fig. 12c. we examined a<br />
total of 21 specimens (10 males and 11 females), all from the northern part of the Gaoligong Shan,<br />
in Bingzhongluo and Cikai Townships, Gongshan County (see Type material above for exact collection<br />
data). These localities are all on the crest or eastern slope of the Gaoligong Shan in Core<br />
Area 2.<br />
OVERALL GEOGRAPHICAL DISTRIBUTION.— This species currently is known only from the<br />
northern part of the Gaoligong Shan, in western Yunnan Province, China.