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DEUVE ET AL.: CARABID BEETLE FAUNA OF THE GAOLIGONG MOUNTAINS 369<br />

Elytra. Elytral silhouette slightly tear-shaped, with humeri effaced, disc markedly convex,<br />

smooth and glabrous. All discal striae effaced except for faintly impressed parascutellar striole and<br />

a short, faintly impressed recurrent stria. Basal setiferous pore present. Three discal setae present,<br />

aligned along presumed track of stria 3. Preapicale seta absent, one or two apicoangular setules<br />

present. Umbilicate setal series with setae of humeral group equidistance from each other, with the<br />

first slightly more medially inserted than the others, setae of median group inserted distinctly posterior<br />

to middle.<br />

Legs. Moderately long but slender. Protibiae with longitudinal furrow, sparely pubescent apically<br />

on anterior surface. Male protarsomeres 1 and 2 dilated and apicomedially toothed.<br />

Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and<br />

ventrite VII of males apically with one pair of paramedial setae, females with two pairs.<br />

Male aedeagus. Median lobe (Fig. 12b) large, elongate, broadest at mid-shaft, with apex long<br />

and recurved dorsally; endophallus with internal sclerites acuminate apically.<br />

COMMENTS.— Based on similarities in the form of the aedeagus of males, this species is closely<br />

related to Queinnectrechus jiuhecola Deuve & Kavanaugh (Deuve et al. 2015), described from<br />

Lijiang County, northwestern Yunnan. However, members of this new species are distinguished<br />

from those of the latter by their much smaller size (BL = 5.3 to 5.8 mm in Q.jiuhecola members),<br />

more elongate body form, more convex elytra, discal striae fully effaced (at least three medial striae<br />

evident in Q.jiuhecola members) and, most significantly, preapical seta absent (present near<br />

stria 2 subapically in Q.jiuhecola members).<br />

In a recent paper, Belousov and Kabak (2016) established a new genus, Uenoites, in which<br />

they placed Q.jiuhecola, as well as three other species previously included in Kozlovites or Deuveotrechus<br />

Uéno (1995), based on the presence of a preapical seta, more convex tempora, and the<br />

median lobe of the male aedeagus not markedly hooked apically. However, all these features are<br />

plesiomorphic among trechines, which is problematic for demonstrating phylogenetic affinity of<br />

the included taxa. Clearly, phylogenetic relationships among these groups of species remain unresolved.<br />

As noted above, the genitalia of Q.griswoldi males are very similar to those of Q.jiuhecola<br />

males, wherease those of the new species described below, Queinnectrechus gongshanicus sp.<br />

nov., are of a very different form, although both of the new species described here are members of<br />

genus Queinnectrechus.<br />

HABITAT DISTRIBUTION.— Members of this species have been found in a variety of microhabitats<br />

in the alpine zone near the crest of the Gaoligong Shan and the eastern slope, at elevations<br />

ranging from 3270 to 3950 m. Specimens were collected under stones in moist meadows and on<br />

tundra slopes and ridges with sparse to thick herbaceous vegetation (Figs. 37b, 38a, 38b, 39b), at<br />

the edges of small streams and seeps from talus slopes, and at the edges of Rhododendron thickets<br />

up to 1.5 m tall. Members of this species have been found syntopic with specimens of Queinnectrechus<br />

(Gaoligongtrechus) balli sp. nov., Queinnectrechus (s.str.) gongshanicus sp. nov., Trechus<br />

gongshanensis sp. nov., Trechus qiqiensis sp. nov. and Trechepaphiopsis monochaeta sp. nov.<br />

GEOGRAPHICAL DISTRIBUTION WITHIN THE GAOLIGONG SHAN.— Fig. 12c. we examined a<br />

total of 21 specimens (10 males and 11 females), all from the northern part of the Gaoligong Shan,<br />

in Bingzhongluo and Cikai Townships, Gongshan County (see Type material above for exact collection<br />

data). These localities are all on the crest or eastern slope of the Gaoligong Shan in Core<br />

Area 2.<br />

OVERALL GEOGRAPHICAL DISTRIBUTION.— This species currently is known only from the<br />

northern part of the Gaoligong Shan, in western Yunnan Province, China.

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