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DEUVE ET AL.: CARABID BEETLE FAUNA OF THE GAOLIGONG MOUNTAINS 399<br />

sagittal aileron, apical lamella short and thick, with apex blunt, endophallus with a sclerotized line<br />

and a scaly area.<br />

DESCRIPTION.— Size medium, BL = 3.7 to 3.8 mm. Body color dark piceous, shiny, elytra<br />

with interval 1, lateral margins and apicomedial area more or less reddish, appendages paler, yellowish-orange,<br />

palpi pale yellow. Body surface smooth, head capsule faintly alutaceous.<br />

Head. Moderate in size; eyes only slightly projected but convex, their diameter twice as long<br />

as tempora, the latter short, moderately convex and glabrous. Frons not flattened; frontal furrows<br />

deep, rounded, prolonged and not attenuated posterior to the eyes. Two pairs of supraorbital setae<br />

present, the anterior pair inserted in foveae. Clypeus with four setae. Labrum with six setae, anterior<br />

margin distinctly concave. Mandibles short; right mandible distinctly tridentate with middle<br />

tooth closer to basal tooth (premolar) than to distal tooth. Mentum and submentum not fused but<br />

nearly so, suture between them only faintly impressed. Mentum with medial tooth apically truncate,<br />

less than half the length of the lateral lobes. Submentum with six setae anteriorly, swollen<br />

anteriorly. Gula wide. Genae with a single seta ventrally on each side. Antennae rather short, with<br />

about 2 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 2 and 3<br />

about equal in length, antennomere 4 slightly shorter.<br />

Pronotum. Transverse (ratio Pw/PL = 1.46), moderately narrowed posteriorly, greatest width<br />

anterior to middle; lateral margins widely rounded, straightened only just anterior to basal angles,<br />

the latter very small, but projected, acute and sharp. Disc convex, smooth and glabrous, median<br />

longitudinal impression slender but sharply impressed, extended anteriorly to near anterior margin;<br />

basal foveae distinct, subcircular; median basal area faintly, longitudinally rugulose, delimited laterally<br />

by obliquely curved furrows. Lateral borders of pronotum moderately slender, narrowly<br />

reflexed, lateral grooves deeply impressed. Single midlateral setae on each side inserted near anterior<br />

one-third; single basolateral seta on each side, inserted at basal angle.<br />

Elytra. Ovoid, only slightly more narrowed basally than apically, humeri distinct but rounded.<br />

Disc convex and smooth. Striae finely impressed, not or only faintly punctate; striae 1 to 3 deeply<br />

impressed on disc, not attenuated apically, remaining striae successively less distinct, the outermost<br />

very faint but still evident. Parascutellar striole present. Recurrent stria terminated anteriorly at distinct<br />

convexity of interval 7. Basal setiferous pore present at common origin of striae 1 and 2. Two<br />

discal setae present and inserted next to stria 3, one at anterior one-fourth and one near middle of<br />

elytra. Preapical seta present and inserted in a prediscal position on interval 3, closer to stria 2 than<br />

3 in forward position at apical one-fourth of elytra. Umbilicate setae of humeral group equidistance<br />

from each other and setae of median group inserted slightly posterior to middle.<br />

Legs. Relatively short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated<br />

and apicomedially toothed.<br />

Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and<br />

ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.<br />

Male aedeagus. Median lobe (Fig. 22b) only moderately enlarged basally but with a mediumsized<br />

sagittal aileron present; shaft narrow subbasally, progressively thicker toward subapical portion;<br />

apical lamella short and thick, with apex blunt, endophallus with a sclerotized line and a scaly<br />

area, similar to that seen in males of T.shiyueliang sp. nov. (Fig. 18b).<br />

HABITAT DISTRIBUTION.— Members of this species have been found under stones on slopes<br />

above (Fig. 43a) and below the road and both side of the pass at an elevation of 3150 m. The habitat<br />

in this area includes broken scrub vegetation of two to four meter high bamboo and Rhododendron<br />

thickets on a thin layer of organic substrate, as well as open areas with stones on granitic sand<br />

substrate, and small seeps. Both specimens were found at the edges of thickets on organic substrate.<br />

No other trechines were found at this site.

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