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426 PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES Series 4, Volume 63, No. 12 area and collected very briefly in “Rhododendron and bamboo thickets” there without success. we concur with him that such thickets represent the most likely habitat for members of this species. Uéno (1999a) reported that specimens of this species (which he described as Trechus fortipes) were found in the same area as specimens of Agonotrechus yunnanus and Trechus indicus (which he recorded as Trechus macrops Jeannel). GEOGRAPHICAL DISTRIBUTION WITHIN THE GAOLIGONG SHAN.— Fig. 32. This species is known only from a single male specimen from the type locality in Baoshan County in the southern part of the Gaoligong Shan region. This locality is in Core Area 7. OVERALL GEOGRAPHICAL DISTRIBU- TION.— This species currently is known only from Baoshan County in the southern part of the Gaoligong Shan region, western Yunnan Province, China. Epaphiotrechus fortipesoides Deuve and Kavanaugh, sp. nov. (Figs. 16g, 33, 46–48) TYPE MATERIAL.— Holotype, a female, in IOZ, labeled: “CASENT 1039089”/ “CHINA, Yunnan, Tengchong County, Houqiao Township, 8.5 airkm NNE of Houqiao at Gaoshidong, 2580 m, N25.39858°/E98.30533°,”/ “27 May 2006, Stop # DHK-2006-043, D.H. Kavanaugh, R.L. Brett, & D.Z. Dong collectors”/ “IMAGE” [green label]/ “HOLOTYPE Epaphiotrechus fortipesoides Deuve and Kavanaugh, sp. nov. designated 2016” [red label]. TYPE LOCALITY.— China, Yunnan, Tengchong County, Houqiao Township, 8.5 airkm NNE of Houqiao at Gaoshidong, N25.39858°/ E98.30533°, 2580 m. DERIVATION OF SPECIES NAME.— The species epithet, fortipesoides, is a combination of the species epithet, fortipes, and the Greek suffix, –ειδής (transliterated into Latin as – oides), meaning resembling, in reference to the similarity of the unique holotype female of this species to members of T.fortipes. FIGURE 32. Epaphiotrechus fortipes (Uéno); Map of locality record (red circle) in the Gaoligong Shan region. Scale line = 100 km.
DEUVE ET AL.: CARABID BEETLE FAUNA OF THE GAOLIGONG MOUNTAINS 427 DIAGNOSIS.— Adults of this species (Fig. 33a) can be distinguished from those of all other species in the region by the following combination of character states: size large (BL = 4.7 mm), apterous; body color piceous, sutural and lateral elytral margins slightly paler, reddish; tempora glabrous; median tooth of mentum apically truncate, almost bifurcate; antennae long; pronotum slightly narrowed (ratio Pw/PL = 1.28), basal angles small, sharp, rectangular; elytra oblong, slightly flattened along median suture, striae finely impressed, finely punctate, striae 1 to 2 or 3 more deeply impressed, more lateral striae more or less effaced but still evident, stria 5 not more deeply impressed at its base; recurrent stria long, deeply impressed, abruptly terminated on interval 7; two discal setiferous pores present, inserted at anterior one-sixth and slightly posterior to middle, respectively, on interval 3 near stria 3; preapical seta present, inserted 1.5 times as far from apical elytral margin as from sutural margin. DESCRIPTION.— Size large, BL = 4.7 mm. Body color dark piceous, head, basal and lateral pronotal margins, and lateral and sutural margins of elytra more or less reddish, appendages paler reddish brown, palpi yellowish brown, dorsum smooth, shiny with pronotum and elytra moderately iridescent, head faintly alutaceous due to moderately impressed isodiametric microsculpture. Head. Broad, slightly elongate. Eyes slightly convex, their diameter slightly less than twice length of tempora, the latter short but distinctly convex. Frons convex, with frontal furrows sharply impressed, not attenuated posterior to eyes. Two pairs of supraorbital setae present, the anterior pair inserted in slight foveae. Clypeus with four setae. Labrum with six setae, anterior margin distinctly concave. Right mandible (Fig. 16g) with premolar not fused with retinaculum but closely associated with the latter, anterior tip of retinaculum enlarged as a distinct tooth and displaced anteriorly. Mentum and submentum not fused. Mentum with medial tooth apically truncate, about onehalf the length of the lateral lobes. Submentum with six setae anteriorly. Genae with a single ventral seta one each side. Antennae moderate in length, with about 2.5 antennomeres extended posteriorly beyond basal pronotal margin; antennomeres 3 and 4 about equal in length, antennomere 2 slightly shorter. Pronotum. Slightly narrowed (ratio Pw/PL = 1.28), with greatest width anterior to middle, only slightly narrowed posteriorly; lateral margins rounded, straightened for a short distance posteriorly, then slightly sinuate just anterior to basal angles, the latter small, sharp and rectangular; basal margin with median region slightly and convexly projected posteriorly. Disc convex, with median longitudinal impression distinct, markedly deepened in median basal area, not extended anteriorly to apical margin; basal foveae moderate in size and depth; median basal area delimited laterally by a pair of short, oblique furrows, smooth except for a pair of paramedial longitudinal foveae. Single midlateral setae on each side inserted anterior to middle; single basolateral seta on each side, inserted at basal angle. Elytra. Elongate ovoid, slightly narrowed anteriorly, humeri rounded and only slightly evident. Disc moderately convex, slightly flattened in sutural area; striae finely impressed, finely punctate, striae 1 to 2 or 3 more deeply impressed, more lateral striae more or less effaced but still evident, stria 5 not more deeply impressed at its base. Parascutellar striole present, deeply impressed and long. Recurrent stria long, deeply impressed, abruptly terminated on interval 7 slightly anterior to the insertion point of preapical seta. Two discal setiferous pores present, inserted at anterior onesixth and near middle, respectively, on interval 3 near stria 3. Preapical seta present, inserted 1.5 times as far from apical elytral margin as from sutural margin on interval 3 next to stria 2. Legs. Short; protibiae with longitudinal furrow. Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and ventrite VII of female apically with two pairs of paramedial setae. COMMENTS.— This species appears to be closely related to E.fortipes, but its members can be
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