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424 PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES<br />

Series 4, Volume 63, No. 12<br />

end abruptly terminated in the posterior end of stria 5. Parascutellar setiferous pore present at base<br />

at common origin of discal striae 1 and 2. Only one discal setiferous pore present, inserted at anterior<br />

two-fifth of elytral length next to stria 3. Only one discal setiferous pore present, inserted<br />

slightly anterior to middle next to stria 3. Preapical seta present, inserted on interval 3 next to stria<br />

2 and about equidistant from apical and sutural elytral margin. Umbilicate setal series with setae of<br />

humeral group equidistance from each other and setae of median group inserted slightly posterior<br />

to middle.<br />

Legs. Short, protibiae with longitudinal furrow. Male protarsomeres 1 and 2 dilated and apicomedially<br />

toothed.<br />

Abdomen. Abdominal ventrites glabrous, except for a single paramedial seta on each side, and<br />

ventrite VII of males apically with one pair of paramedial setae, of females with two pairs.<br />

Male aedeagus. Median lobe (Fig. 31b) with sagittal aileron very small, shaft moderately<br />

broad basally, then gradually narrowed apically to a narrowly rounded apex; endophallus with a<br />

faintly sclerotized scaly area distinctly elongate and narrowed distally.<br />

COMMENTS.— This species appears to be very closely related (perhaps sister species) to<br />

T.monochaetus, but its male members can be distinguished from those of the latter by having the<br />

base of the median lobe of the male aedeagus narrower (broader in T.monochaeta members, see<br />

Fig. 30b) and the scaly sclerotized area of the endophallus distinctly narrowed distally (of more<br />

equal width throughout in T.monochaeta males).<br />

HABITAT DISTRIBUTION.— Members of this species have been collected by sifting leaf litter<br />

from forests of scattered, large Abies sp. trees with a dense understory of bamboo or Rhododendron<br />

spp. at elevations ranging from 3221 to 3290 m. Specimens of Trechus shiyueliang andT.<br />

shibalicus were collected in one or more of the same litter samples. Specimens of T.unipilosa were<br />

also collected at 3100 m elevation under stones at the open edges of a small stream draining a<br />

north-facing glacial cirque with a large snowfield in its basin (Fig. 40b). A specimen of Trechepaphiopsis<br />

unisetulosa sp. nov. was also collected in this area, at the upper limit of the altitudinal<br />

range of this last mentioned species.<br />

GEOGRAPHICAL DISTRIBUTION WITHIN THE GAOLIGONG SHAN.— Fig. 31c. we examined a<br />

total of 18 specimens (13 males and 5 females) from sites on the eastern slope of the northcentral<br />

part of the Gaoligong Shan in Fugong County (see Type material above for exact collection data).<br />

All of these sites are in Core Area 3.<br />

OVERALL GEOGRAPHICAL DISTRIBUTION.— This species currently is known only from Fugong<br />

County in the northcentral part of the Gaoligong Shan region, western Yunnan Province, China.<br />

Genus Epaphiotrechus Deuve and Kavanaugh, gen. nov.<br />

TYPE SPECIES.— Epaphiotrechus fortipesoides sp. nov.<br />

Derivation of genus group name.— The genus group name (masculine) is a combination of<br />

two other trechine generic names, Epaphius and Trechus.<br />

DIAGNOSIS.— Adults of this genus (Fig. 33a) can be recognized by the following combination<br />

of character states: size large (BL = 4.5 to 4.7 mm), apterous; body color dark piceous with elytral<br />

interval 1 and lateral areas of pronotum and elytra more or less reddish; dorsum shiny, distinctly<br />

iridescent, dorsal surface glabrous except for isolated fixed setae typical for trechines, eyes also<br />

glabrous; right mandible (Fig. 16g) with premolar not fused with retinaculum but closely associated<br />

with the latter [possibly representative of an intermediate state in the evolution of the “bidentate”<br />

mandibular type (see discussion above for genus Trechus)], anterior tip of retinaculum<br />

enlarged as a distinct tooth and displaced anteriorly (but not quite as far as in members of Trechepaphiopsis<br />

species); pronotum with basal angles small and rectangular; elytra elongate, oblong,

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