360 PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES Series 4, Volume 63, No. 12 FIGURE 9. Agonotrechus xiaoheishan sp. nov.; a. Dorsal habitus (CASENT1036866). b. Median lobe of aedeagus of male (CASENT1036866), left lateral aspect. c. Map of locality records (red circles) for A.xiaoheishan in the Gaoligong Shan region. Scale lines a, b = 0.5 mm, c = 100 km.
DEUVE ET AL.: CARABID BEETLE FAUNA OF THE GAOLIGONG MOUNTAINS 361 eral lobes. Submentum with six setae subapically, gula broad. Genae with a single seta ventrally on each side. Antennae long and slender, extended to (in females) or slightly beyond (in males) the middle of the elytra, with five or six antennomeres beyond the pronotal base; antennonmeres 3 and 4 virtually the same length and antennomere 2 slightly shorter. Pronotum. Relatively small, not or only slightly narrowed basally, only slightly transverse, ratio Pw/PL = 1.21, widest at anterior one-third; lateral margins rectilinear in basal half, not sinuate except for a very slight and short inflexion just anterior to the basal angle, which is subrectangular and sharp; pronotal disc smooth, glabrous, moderately convex; median longitudinal impression very fine; basal foveae deep, basal flattened area rather small, delimited laterally by short but deep and oblique furrows. Lateral explanation slender and moderately reflexed in anterior twothirds, then progressively broader in basal one-third without. Midlateral pair of setae inserted at anterior one-fifth and basolateral pair inserted at hind angles. Elytra. Convex, large and broad, especially in relation to pronotum, elytral silhouette ovoid, about equally narrowed apically and basally, humeri evident but rounded. All discal striae evident, regular, deeply impressed and punctate, however striae 1 to 4 effaced or nearly so near base, parascutellar striole rather long, recurrent stria continuous anteriorly with discal stria 5, intervals convex, only a single discal seta (anterior seta) present, inserted at basal one-eighth near stria 3. Parascutellar setiferous pore present at base at common origin of discal striae 1 and 2. Anterior discal seta present, inserted at basal one-eighth of elytra in stria 3, which is effaced anterior to that point. Middle discal seta absent. Preapical seta present on interval 3 near stria 2 opposite the anterior edge of the subapical sinuation. Umbilicate setal series with setae of humeral group equidistant for each other and those of median group both inserted posterior to middle of elytra. Legs. Slender but only moderately long. Protibiae furrowed. Male protarsi with tarsomeres 1 and 2 dilated and apicomedially toothed. Male aedeagus. Median lobe (Fig. 9b) short, with apex bent ventrally and bluntly pointed, endophallus with a spoon-shaped sclerite. COMMENTS.— Members of this new species are morphologically similar to those of Agonotrechus birmanicus Bates (1892), described from Kachin State in Myanmar, and Agonotrechus tenuicollis Uéno (1986), described from eastern Nepal. However, they can be distinguished from members of both of these other species by the following features: pronotum with anterior margin straight, not concave, anterior angles only faintly projected and more broadly rounded, and lateral borders more slender anteriorly and at middle; and apex of the median lobe of the male aedeagus narrower and more curved. HABITAT DISTRIBUTION.— Members of this species have been found in and under rotting logs in dark, closed-canopy broadleaf evergreen forest (Fig. 44) at elevations ranging from 2020 to 2067 m. Although members of no other Agonotrechus species have been found syntopic with those of A. xiaoheishan, specimens of Trechus indicus were collected in the same samples. GEOGRAPHICAL DISTRIBUTION WITHIN THE GAOLIGONG SHAN.— Fig. 9c. we examined a total of 14 specimens (4 males and 10 females), all from Xiaoheishan Forest Reserve in the southern part of the Gaoligong Shan (see Type material above for exact collection data). This species was recorded only from near the top of the western slope in the southern part of the study area (Core Area 6). Its known geographical range does not overlap with that of any other Agonotrechus species, although A.yunnanus has been recorded from the adjacent Core Area 7 on the eastern slope of the Gaoligong Shan, 13.3 km to the north. OVERALL GEOGRAPHICAL DISTRIBUTION.— This species currently is known only from the type area in the southern part of the Gaoligong Shan, in western Yunnan Province, China.
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