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GUNS, GERMS AND STEEL - Cloverport Independent Schools

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APPLES OR INDIANS • 1 3 7<br />

tional changes had to be made in them under cultivation. As we discussed<br />

in the preceding chapter, the principal changes—the breakdown of the<br />

natural systems of seed dispersal and of germination inhibition—evolved<br />

automatically and quickly as soon as humans began to cultivate the seeds<br />

in fields. The wild ancestors of our wheat and barley crops look so similar<br />

to the crops themselves that the identity of the ancestor has never been in<br />

doubt. Because of this ease of domestication, big-seeded annuals were the<br />

first, or among the first, crops developed not only in the Fertile Crescent<br />

but also in China and the Sahel.<br />

Contrast this quick evolution of wheat and barley with the story of<br />

corn, the leading cereal crop of the New World. Corn's probable ancestor,<br />

a wild plant known as teosinte, looks so different from corn in its seed and<br />

flower structures that even its role as ancestor has been hotly debated by<br />

botanists for a long time. Teosinte's value as food would not have<br />

impressed hunter-gatherers: it was less productive in the wild than wild<br />

wheat, it produced much less seed than did the corn eventually developed<br />

from it, and it enclosed its seeds in inedible hard coverings. For teosinte to<br />

become a useful crop, it had to undergo drastic changes in its reproductive<br />

biology, to increase greatly its investment in seeds, and to lose those rocklike<br />

coverings of its seeds. Archaeologists are still vigorously debating how<br />

many centuries or millennia of crop development in the Americas were<br />

required for ancient corn cobs to progress from a tiny size up to the size<br />

of a human thumb, but it seems clear that several thousand more years<br />

were then required for them to reach modern sizes. That contrast between<br />

the immediate virtues of wheat and barley and the difficulties posed by<br />

teosinte may have been a significant factor in the differing developments<br />

of New World and Eurasian human societies.<br />

A third advantage of the Fertile Crescent flora is that it includes a high<br />

percentage of hermaphroditic "selfers"—that is, plants that usually pollinate<br />

themselves but that are occasionally cross-pollinated. Recall that most<br />

wild plants either are regularly cross-pollinated hermaphrodites or consist<br />

of separate male and female individuals that inevitably depend on another<br />

individual for pollination. Those facts of reproductive biology vexed early<br />

farmers, because, as soon as they had located a productive mutant plant,<br />

its offspring would cross-breed with other plant individuals and thereby<br />

lose their inherited advantage. As a result, most crops belong to the small<br />

percentage of wild plants that either are hermaphrodites usually pollinating<br />

themselves or else reproduce without sex by propagating vegetatively

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