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In Pursuit of the Gene

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52 ¨ GALTON’S DISCIPLES<br />

bly is continuous or nearly so.” What he meant to argue was that “in <strong>the</strong><br />

case <strong>of</strong> particular teeth evidence <strong>of</strong> discontinuous variation has been given.”<br />

The problem has been fur<strong>the</strong>r compounded, he now realized, by his failure<br />

to give a more precise definition <strong>of</strong> discontinuity, Bateson feared, so that<br />

“each hostile reader remains free to attribute to it <strong>the</strong> one particular meaning<br />

which strikes him as especially foolish and that he can most easily show<br />

to be untrue.” 18<br />

Two days later, Weldon hastened to assure Bateson that he had not approached<br />

Bateson’s book with “any unduly hostile spirit,” and yet, he<br />

added, “it does not even yet seem to me that you make a clear case for discontinuity<br />

in any single instance.” Weldon proceeded to eviscerate Bateson’s<br />

argument with elegant prose, much <strong>of</strong> which he included nearly verbatim<br />

in a Nature review that appeared in May. While acknowledging <strong>the</strong> vast effort<br />

that Bateson had put into collecting evidence for his book, Weldon argued<br />

that Bateson had failed to make a compelling case for discontinuous<br />

evolution. The concluding paragraph <strong>of</strong> Weldon’s review, which was a masterpiece<br />

<strong>of</strong> understatement, cast serious doubts about <strong>the</strong> significance <strong>of</strong><br />

Bateson’s seven-year 19 effort to document <strong>the</strong> existence <strong>of</strong> discontinuous<br />

variation:<br />

The only way in which <strong>the</strong> question can be settled for a given variation<br />

seems to be by taking large numbers <strong>of</strong> animals, in which <strong>the</strong> variation is<br />

known to occur, at random, and making a careful examination and record<br />

<strong>of</strong> each. Mr. Bateson’s chapter on teeth, like all his chapters, is <strong>of</strong><br />

great interest, and will doubtless serve to throw important light on many<br />

things. But a careful histological account <strong>of</strong> 500 dogs would have done<br />

more to show <strong>the</strong> least possible size <strong>of</strong> a tooth in dogs than all <strong>the</strong> information<br />

so painfully collected. And so in many o<strong>the</strong>r cases. 20<br />

Weldon’s negative opinion <strong>of</strong> Bateson’s book was widely shared by <strong>the</strong><br />

scientific community, which was still in <strong>the</strong> thrall <strong>of</strong> <strong>the</strong> Darwinian orthodoxy<br />

<strong>of</strong> evolution by incremental steps. Pr<strong>of</strong>essors did not recommend <strong>the</strong><br />

book to be read by <strong>the</strong>ir students, and it was soon remaindered. <strong>In</strong> <strong>the</strong> darkness<br />

<strong>of</strong> <strong>the</strong> dismal reception, <strong>the</strong>re was one ray <strong>of</strong> light: Francis Galton<br />

loved <strong>the</strong> book. “It was,” Galton wrote in <strong>the</strong> July issue <strong>of</strong> <strong>the</strong> widely read

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