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Principios de Taxonomia

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4.2 What Basic Rule Defines Traits as Being Taxonomically Relevant?j69<br />

traits that distinguish every particular organism from others in a population. Every<br />

organism is unique in its complexity.<br />

Every biologist is aware of this phenomenon. However, less commonly known is<br />

the obvious problem of how one can actually speak of species differences if the<br />

members of a species differ so drastically in regard to their traits. What is the actual<br />

difference in traits between species when all organisms within a species differ from<br />

each other? Only monozygotic twins are i<strong>de</strong>ntical in traits.<br />

The phenomenon of allelic diversity conjures the difficult question of what are<br />

intraspecific differences in contrast to interspecific differences. What is behind the<br />

fact that different species are distinguished from each other by their traits? Every user<br />

of an animal or plant i<strong>de</strong>ntification gui<strong>de</strong> uses the option of distinguishing one<br />

species from another species by certain traits. Is it the extent of differences that makes<br />

it possible to distinguish species, such as the principle that few differences equals<br />

belonging to the same species but many differences equals belonging to a different<br />

species? Or is it the quality of differences that makes it possible to distinguish species,<br />

such as the principle that certain differences equals belonging to the same species<br />

and other differences equals belonging to a different species?<br />

Neither is the case. Numeric criterion can be eliminated immediately. It cannot be<br />

the extent of differences. Otherwise, males and females from a great number of<br />

animal species would have to belong to different species. Qualitative criteria can also<br />

be refused. The principle of What only differs in the number of bristles belongs to<br />

one and the same species, but what differs in the number of legs are different<br />

species, cannot be generalized either.<br />

Every multicellular organism has many thousands of traits. Even today in the age of<br />

genome sequencing, in which the number of animal and plant species whose<br />

genomes are completely sequenced continually grows, there is no un<strong>de</strong>rstanding<br />

of how many phenotypic traits a particular species possesses in contrast to another<br />

species. The logical conclusion is that the characterization of a species by its traits<br />

always has to be based on a restricted selection of traits and that all traits can never be<br />

consi<strong>de</strong>red. We presumably will be able to do this one day after the sequencing of all<br />

genes and all gene regulatory and synthesis pathways have been <strong>de</strong>ciphered.<br />

However, this day still lies in the far distant future. Currently, every taxonomical<br />

classification is assigned according to traits that are based on the selection of a few<br />

traits. This selection is subjective.<br />

The scientist, however, should have the <strong>de</strong>sire to establish an un<strong>de</strong>rlying system of<br />

rules for general validity, which <strong>de</strong>fine the quality of traits suitable for taxon<br />

membership. Such a theoretical foundation for a trait-based taxonomy should <strong>de</strong>fine<br />

for all organisms a system of rules for which traits are relevant for taxonomic<br />

classification and which are not. However, this <strong>de</strong>sire cannot be fulfilled.<br />

The traits that distinguish one species from another are not quantifiable. It is not<br />

possible to <strong>de</strong>termine a number of traits that are species specific. From this, it follows<br />

that a statement such as the following is senseless: The organisms of two different<br />

populations match with regard to half of their traits. For example, Wilhelm Meise<br />

claimed 70 years ago that the Italian Sparrow (Passer italiae) was a hybrid species of<br />

the House Sparrow (Passer domesticus) and the Spanish Sparrow (Passer hispaniolensis)

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