Principios de Taxonomia
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200j 7 The Cohesion of Organisms Through Genealogical Lineage (Cladistics)<br />
reasons why paraphyly cannot be eradicated. What justification is there for a<br />
paraphyletic taxonomy? There are only pragmatic reasons for this; there is no<br />
consistent un<strong>de</strong>rlying theory.<br />
Only the human <strong>de</strong>sire to combine organisms into a common group that look<br />
similar to the human eye can motivate the use of paraphyletic groups (Figure 2.4:<br />
B þ C is a paraphyletic group). This is not at all a scientific line of thinking. The<br />
formation of paraphyletic groups means that two different classification principles<br />
are intermixed: grouping according to the <strong>de</strong>scent relation and classification according<br />
to trait resemblance.<br />
The best-known example of paraphyly in biology is the aforementioned class of<br />
reptiles along with the class of birds within the phylum of vertebrates (Figure 7.6). The<br />
reptiles are an incomplete, truncated taxon because the birds that branch off from<br />
the rest of the reptiles at an evolutionarily later time have been withdrawn from the<br />
reptile class. However, the birds are the sisters of the dinosaurs and are closely related to<br />
these, while both together (the birds and the dinosaurs) are less closely related to the<br />
crocodiles, snakes and turtles than birds and dinosaurs are to each other.<br />
Thus, the formation of the class of reptiles without the birds contradicts the logic of<br />
a kinship-oriented taxonomy. This is not taxonomy according to kinship, but<br />
taxonomy according to the assessment of traits. However, the assessment of traits<br />
is based on subjective standards that are set by humans ourselves. This means that<br />
paraphyla (in contrast to monophyla) are artificial groups, whose coherence and<br />
boundaries as separate groups are not <strong>de</strong>monstrated in nature (Figure 2.4: B þ C is a<br />
paraphyletic group). For the empirically driven scientist who wants to restrict himself<br />
to phenomena that can be objectively observed in nature, paraphyly is a group<br />
formation that is highly questionable, if not to be rejected altogether.<br />
We humans think that birds simply look very different from the reptiles, but we are<br />
unable to quantify this distinction. Birds are warm-bloo<strong>de</strong>d, have feathers, are usually<br />
multicolored and often sing in a manner pleasant to the human ear. Accordingly, in<br />
the eyes of humans, they distinguish themselves drastically from crocodiles, snakes<br />
and turtles. These are cold-bloo<strong>de</strong>d, have scales or horny scutes, are usually colored<br />
inconspicuously, and make utterances that are almost without exception unpleasant<br />
to humans. The qualities of these criteria cannot be reason to group birds and reptiles<br />
into separate equal-ranked taxa.<br />
For the mo<strong>de</strong>rn, factually driven scientist, it is disconcerting that these kinds of<br />
criteria serve as a reason to separate the birds from the reptiles and award them an<br />
equal rank along with the rest of the reptiles. Why do the possession of a plumage and<br />
warm-bloo<strong>de</strong>dness rank so high that the birds are ascribed such dissimilarity to the<br />
remaining reptiles? By reverting less to subjective human perception and instead<br />
perhaps valuing certain metabolic enzymes that are not visible to the human eye, the<br />
birds would not be such a unique case compared to the rest of the reptiles. Then, the<br />
birds autonomy would be en<strong>de</strong>d, and the birds would unambiguously belong to<br />
the reptiles, with which they are closely related and from which they should not be<br />
segregated.<br />
Let the contrariness of a trait-based and a kinship-based grouping once again be<br />
<strong>de</strong>monstrated by the following example (Figure 7.7). Of the four grandchildren