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Principios de Taxonomia

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2.14 The Dualism of the Species Concept: the Epistemic vs. the Operative Goalj41<br />

<strong>de</strong>fining daytime by the trait of brightness is wrong. The <strong>de</strong>finition is wrong because<br />

daytime is not what brightness specifies.<br />

Correctly, daytime (as opposed to nighttime) is <strong>de</strong>fined by the position of Earth<br />

relative to the sun, not by its brightness. Daytime is the period of time at which the<br />

surface of the Earth is turned towards the sun. Nighttime is the period of time at<br />

which the surface of the Earth is turned away from the sun. In contrast, brightness is<br />

only a symptom of daytime, and darkness is a symptom of nighttime. These<br />

<strong>de</strong>finitions are operational <strong>de</strong>finitions. Bright light can be present even at nighttime.<br />

However, it can never be daytime in one location if the sun is positioned on the other<br />

si<strong>de</strong> of the Earth. Summer and winter on Earth are <strong>de</strong>fined by the inclination of the<br />

Earth s axis, not by their traits of being warm or cold.<br />

Exactly the same type of reasoning applies to the biological species. The goal of<br />

i<strong>de</strong>ntifying a given individual as a member of a given species should never be<br />

confused with the goal of knowing that a given group of organisms is a species.<br />

Nothing expresses the difference between diagnostics and ontology better than a<br />

statement by the classical author of evolutionary theory, American paleontologist<br />

George Gaylord Simpson (Simpson, 1961): The <strong>de</strong>finition of monozygotic twins<br />

... provi<strong>de</strong>s a homologous causal sequence. ... Two monozygotic twins are<br />

not twins because they are similar but, quite the contrary, are similar because<br />

they are twins.<br />

This quote expresses in a direct manner that on one level, there concern is about<br />

i<strong>de</strong>ntifying twins. On another level, the quote addresses a markedly different issue,<br />

which is that twins are twins because they originate from a single egg. Twins have<br />

i<strong>de</strong>ntical traits as a result of being twins, but this is not the ontological <strong>de</strong>finition of<br />

twins because even non-twins can have i<strong>de</strong>ntical traits. The fact that there are many<br />

field gui<strong>de</strong>s on the i<strong>de</strong>ntification of species on the market, all of which focus on<br />

species diagnostics, may mislead us into falsely believing that diagnostic differences<br />

are ontological differences.<br />

2.14<br />

The Dualism of the Species Concept: the Epistemic vs. the Operative Goal<br />

How can we finally solve the species problem, meaning the worldwi<strong>de</strong> dissension<br />

about what a species is? Different authors mean different things by the word<br />

species (Mallet, 1995). For more than a hundred years, this has led to what is<br />

called the species problem . Disagreements about what a species is have led to <strong>de</strong>ep<br />

dissent, so much so that it is referred to as a never-ending story.<br />

Taxonomists place varying weights on trait differences between organisms,<br />

<strong>de</strong>scent or sexual cohesion between organisms. Why are there approximately<br />

9000 bird species on Earth (<strong>de</strong>l Hoyo, Elliott, and Sargatal, 1992)? It would be easy<br />

and consistent to <strong>de</strong>fend other estimates, resulting in as many as 27 000 species<br />

(Cracraft, 1997). The number <strong>de</strong>pends entirely on the weight given to particular<br />

<strong>de</strong>limiting criteria. There are many diagnostically easily recognizable subgroups,<br />

distinguishable races, genetically far-distanced groups or allopatrically isolated

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