Principios de Taxonomia
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2.14 The Dualism of the Species Concept: the Epistemic vs. the Operative Goalj41<br />
<strong>de</strong>fining daytime by the trait of brightness is wrong. The <strong>de</strong>finition is wrong because<br />
daytime is not what brightness specifies.<br />
Correctly, daytime (as opposed to nighttime) is <strong>de</strong>fined by the position of Earth<br />
relative to the sun, not by its brightness. Daytime is the period of time at which the<br />
surface of the Earth is turned towards the sun. Nighttime is the period of time at<br />
which the surface of the Earth is turned away from the sun. In contrast, brightness is<br />
only a symptom of daytime, and darkness is a symptom of nighttime. These<br />
<strong>de</strong>finitions are operational <strong>de</strong>finitions. Bright light can be present even at nighttime.<br />
However, it can never be daytime in one location if the sun is positioned on the other<br />
si<strong>de</strong> of the Earth. Summer and winter on Earth are <strong>de</strong>fined by the inclination of the<br />
Earth s axis, not by their traits of being warm or cold.<br />
Exactly the same type of reasoning applies to the biological species. The goal of<br />
i<strong>de</strong>ntifying a given individual as a member of a given species should never be<br />
confused with the goal of knowing that a given group of organisms is a species.<br />
Nothing expresses the difference between diagnostics and ontology better than a<br />
statement by the classical author of evolutionary theory, American paleontologist<br />
George Gaylord Simpson (Simpson, 1961): The <strong>de</strong>finition of monozygotic twins<br />
... provi<strong>de</strong>s a homologous causal sequence. ... Two monozygotic twins are<br />
not twins because they are similar but, quite the contrary, are similar because<br />
they are twins.<br />
This quote expresses in a direct manner that on one level, there concern is about<br />
i<strong>de</strong>ntifying twins. On another level, the quote addresses a markedly different issue,<br />
which is that twins are twins because they originate from a single egg. Twins have<br />
i<strong>de</strong>ntical traits as a result of being twins, but this is not the ontological <strong>de</strong>finition of<br />
twins because even non-twins can have i<strong>de</strong>ntical traits. The fact that there are many<br />
field gui<strong>de</strong>s on the i<strong>de</strong>ntification of species on the market, all of which focus on<br />
species diagnostics, may mislead us into falsely believing that diagnostic differences<br />
are ontological differences.<br />
2.14<br />
The Dualism of the Species Concept: the Epistemic vs. the Operative Goal<br />
How can we finally solve the species problem, meaning the worldwi<strong>de</strong> dissension<br />
about what a species is? Different authors mean different things by the word<br />
species (Mallet, 1995). For more than a hundred years, this has led to what is<br />
called the species problem . Disagreements about what a species is have led to <strong>de</strong>ep<br />
dissent, so much so that it is referred to as a never-ending story.<br />
Taxonomists place varying weights on trait differences between organisms,<br />
<strong>de</strong>scent or sexual cohesion between organisms. Why are there approximately<br />
9000 bird species on Earth (<strong>de</strong>l Hoyo, Elliott, and Sargatal, 1992)? It would be easy<br />
and consistent to <strong>de</strong>fend other estimates, resulting in as many as 27 000 species<br />
(Cracraft, 1997). The number <strong>de</strong>pends entirely on the weight given to particular<br />
<strong>de</strong>limiting criteria. There are many diagnostically easily recognizable subgroups,<br />
distinguishable races, genetically far-distanced groups or allopatrically isolated