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Conspectus cobitidum - Raffles Museum of Biodiversity Research

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locality: Kazakhstan: Alma-Atinskaya oblast: Kaskelen<br />

River, 2 km before it enters Ili River; holotype: ZMMU<br />

P-3638; noun in genitive, indeclinable)<br />

10.42.93 Triplophysa shaanxiensis Chen, in Chen, Xu,<br />

Fang, Song & Wang, 1987<br />

Triplophysa shaanxiensis Chen, in Chen, Xu, Fang, Song &<br />

Wang, 1987: 21, fig. 8 (type locality: China: Shaanxi:<br />

Tongchuan Xian; holotype: SIZX 801811; adjective, -is,<br />

-is, -e)<br />

10.42.94 Triplophysa shehensis Menon, 1987<br />

Triplophysa shehensis Menon, 1987: 212, pl. 16 fig. 9 (type<br />

locality: India: Kashmir: Ladakh: Sheh, irrigation canal,<br />

nearly 12 km east <strong>of</strong> Leh; holotype: ZSI/NRS uncat.;<br />

adjective, -is, -is, -e)<br />

Noemacheilus shehensis Tilak, 1990: 138, figs. 2–3 (type<br />

locality: India: Kashmir: Ladakh: irrigation canal at Sheh,<br />

nearly 12 km east <strong>of</strong> Leh; holotype: ZSI/NRS 1208; junior<br />

secondary homonym <strong>of</strong> Triplophysa shehensis Menon,<br />

1987: 212; paper dated 1990 but possibly not published<br />

until 1997; adjective, -is, -is, -e)<br />

Nomenclatural notes. Menon (1987) described this species<br />

and indicated the author as Tilak, then in press. Tilak's paper<br />

only appeared in 1990 (date on the cover, actual publication<br />

possibly much later, the paper is first listed in Zoological<br />

Record for 1997). The description in Menon is based on a<br />

single <strong>of</strong> Tilak's specimens and is not extracted from Tilak's<br />

text. Thus Menon is author <strong>of</strong> the name and the single specimen<br />

he examined is the holotype. The locality data listed<br />

by Menon is only "irrigation canal, Sheh" but as all <strong>of</strong> Tilak's<br />

(1997) specimens are from a single locality, it can be<br />

completed as above. Tilak's N. shehensis, being based on a<br />

separate description, without reference to Menon's description,<br />

is a distinct nominal species and a junior secondary<br />

homonym <strong>of</strong> T. shehensis when transferred to Triplophysa.<br />

It is not known whether the holotype <strong>of</strong> the two nominal<br />

species is the same specimen or not, which would make them<br />

objective synonyms.<br />

10.42.95 Triplophysa shilinensis Chen & Yang, in Chen,<br />

Yang & Xu, 1992<br />

Triplophysa shilinensis Chen & Yang, in Chen, Yang & Xu,<br />

1992: 17, fig. 1 (type locality: China: Yunnan: Lunan<br />

County: cave at Weiboyi village near Shi Lin Stone Forest<br />

[24°47'N 103°22'E; Romero et al., 2009: 263]; holotype:<br />

KIZ 913001; adjective, -is, -is, -e)<br />

10.42.96 Triplophysa shiyangensis (Zhao & Wang, 1983)<br />

Nemachilus shiyangensis Zhao & Wang, 1983: 97, fig. 1<br />

(type locality: China: Gansu: Leitai He stream near Wuwei<br />

[37°55'N 102°38'E], Shiyang He River system, "Huang<br />

He drainage" [in fact, Shiyang is an endorheic river<br />

ending in Gobi desert about 39°00'N 103°35'E]; holotype:<br />

DBLU 770582; adjective, -is, -is, -e)<br />

10.42.97 Triplophysa siluroides (Herzenstein, 1888)<br />

Nemachilus siluroides Herzenstein, 1888: 62, pl. 7 fig. 1,<br />

pl. 8 fig. 10 (type locality: China: Qinghai: Chuan-che<br />

River near Gomi [Huang He River near Balekhun Gomi,<br />

THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />

131<br />

approx. 30°10'N 100°45'E; now flooded in Longyangxia<br />

Reservoir]; syntypes: ZISP 7243 [1], 7244 [1], 7851 [1])<br />

10.42.98 Triplophysa stenura (Herzenstein, 1888)<br />

Nemachilus stenurus Herzenstein, 1888: 64, pl. 1 fig. 1 (type<br />

locality: China: Tibet: Dy-Tschu [Tongtian He, in upper<br />

Yangtze basin; He et al., 2008: 48]; lectotype ZISP 7355,<br />

designated by Prok<strong>of</strong>iev, 2009a: 699; compound adjective,<br />

-us, -a, -um)<br />

? Nemachilus lhasae Regan, 1905b: 301 (type locality: Tibet:<br />

Lhasa; holotype: BMNH 1905.2.8.16; noun in genitive,<br />

indeclinable)<br />

Taxonomic notes. Triplophysa stenura is presently reported<br />

from the upper Yangtze, Mekong, Salween and Brahmaputra<br />

drainages, which suggests that several species are confused<br />

under this name.<br />

10.42.99 Triplophysa stewarti (Hora, 1922)<br />

Diplophysa stewarti Hora, 1922: 70, fig. 2c–d (type locality:<br />

China: Xizang: Rham-tso [Hram Tso, Bam Tso;<br />

28°08'52"N 89°20'02"E]; holotype: ZSI F 2894/1; noun<br />

in genitive, indeclinable)<br />

Nemachilus panguri Hora, 1936: 318, figs. 8–9, pl. 12 figs.<br />

3–4 (type locality: India: Jammu & Kashmir: Ladakh:<br />

Pangur Tso [Spanggur Tso, 33°32'N 78°55'E] / China:<br />

Xizang: Tso Nyak [apparently at 33°29'28"N 80°18'11"E]<br />

[both in Pangong Tso endorheic basin]; syntypes ["several"]:<br />

ZSI F 12204/1 [2], Menon & Yazdani, 1968: 123;<br />

noun in genitive, indeclinable)<br />

Nemachilus longianalis Ren & Wu, 1982: 82, fig. 2 (type<br />

locality: China: Northern Xizang: Nam Cuo Lake [? Nam<br />

Co, Nam Tso, 30°42'N 90°33'E]; holotype: HFRI 80-<br />

1669; compound noun, indeclinable)<br />

Nemachilus longianalis tangtianhensis Wu & Wu, 1988a:<br />

21, fig. 4 (type locality: China: Qinghai: mouth <strong>of</strong> Dangqu,<br />

upper reaches <strong>of</strong> Tangtianhe River; holotype: NPIB<br />

8606088; adjective, -is, -is, -e)<br />

10.42.100 Triplophysa stolickai (Steindachner, 1866)<br />

Cobitis Stoličkai Steindachner, 1866: 793, pl. 14 fig. 2 (type<br />

locality: India: Ladakh: Rupshu: rivulets in vicinity <strong>of</strong><br />

Lake Tsumureri [Tso Morari] [contra Eschmeyer &<br />

Fricke, 2010, "Umgebung" is not the name <strong>of</strong> a river but<br />

the German word meaning vicinity]; syntypes [12]: NMW<br />

48436 [5], 48439 [1], 50477 [4], Eschmeyer & Fricke,<br />

2010; the č in original spelling must be emended in c,<br />

Code art. 32.5.2.1; noun in genitive, indeclinable)<br />

Taxonomic notes. Prok<strong>of</strong>iev (2007b: 4) recognised T. stolickai<br />

with a number <strong>of</strong> subspecies. I do not recognise these<br />

subspecies because the Evolutionary Species Concept used<br />

here does not accomodate subspecies, and because these 'subspecies'<br />

are diagnosable lineages occupying very distinct and<br />

isolated areas and fulfill the criteria to be recognised as species.<br />

See T. cakaensis, T. dorsonotata, T. hexiensis, T. obtusirostra,<br />

T. retropinnis and T. uranoscopus.<br />

10.42.101 Triplophysa strauchii (Kessler, 1874)<br />

Diplophysa Strauchii Kessler, 1874: 58, pl. 8 fig. 40 (type<br />

locality: Kazakhstan: Ili River, tributary <strong>of</strong> Lake Balkash /<br />

Ich-Balya River; syntypes: ZISP 2328 [2], 2329 [1],

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