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Conspectus cobitidum - Raffles Museum of Biodiversity Research

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(type locality: India: Manipur: Chindwin drainage,<br />

Lokchao River near Moreh, 110 km from Imphal City;<br />

holotype: MUMF 3333/1A; adjective, -is, -is, -e)<br />

Nomenclatural notes. Words ending in -cola and meaning<br />

'inhabitant <strong>of</strong>' are nouns and indeclinable.<br />

7.7.4 Homalopteroides smithi (Hora, 1932)<br />

Homaloptera smithi Hora, 1932a: 286, pl. 11 fig. 3 (type<br />

locality: Thailand: Nakon Sritamarat Province: Tadi<br />

Stream and Klong Pong at Ban Kiriwong; syntypes:<br />

KUMF 165 [3], USNM 107941 [1], 109821 [5],<br />

119459 [1], ZSI F 11293–11294/1 [4], 11295/1 [2], Menon<br />

& Yazdani, 1968: 118, Eschmeyer & Fricke, 2010;<br />

noun in genitive, indeclinable)<br />

? Homaloptera maxinae Fowler, 1937: 152, figs. 52–53 (type<br />

locality: Thailand: Tachin [Tha Chin, Samut Sakhon;<br />

13°32'22"N 100°15'20"E]; holotype: ANSP 68004; noun<br />

in genitive, indeclinable)<br />

Homaloptera lineata Smith, 1945: 277, fig. 55 (type locality:<br />

Thailand: Mekong at Chiangsen Kao; holotype:<br />

USNM 199488; adjective, -us, -a, -um)<br />

? Homaloptera indochinensis Silas, 1953: 192, fig. 2 (type<br />

locality: "Indo-China (? Tonkin)" [Vietnam: Kontum; see<br />

below]; holotype: BMNH 1933.8.19.50; adjective, -is,<br />

-is, -e)<br />

Taxonomic notes. Homaloptera indochinensis is based on<br />

a single specimen without precise locality. From Silas's description,<br />

it appears to fall within the variation <strong>of</strong> H. smithi,<br />

a very common species in the Mekong drainage. From information<br />

by E. Trewavas to S. L. Hora (in Silas, 1951: 11),<br />

the specimen had no locality label but Delacour & Lowe<br />

had collected in "Pekh, N. Tonkin [northern Vietnam; ? Pakha];<br />

Xien Khonang or Chin chuang, Laos [Xieng Khouang]<br />

and Kontum and Dak-to in Annam [Vietnam: Kontum and<br />

Dak To]". This material is registered as BMNH 1933.8.19.1–<br />

100. In fact this material had already been catalogued earlier<br />

as BMNH 1933.8.4.1–100. In this earlier list this specimen<br />

appears as H. wassinki, BMNH 1933.8.4.51, and the<br />

locality is listed as Kontum.<br />

Delacour conducted six expeditions to Laos, Cambodia<br />

and Vietnam between 1923 and 1939; Lowe accompanied<br />

him in five <strong>of</strong> these. There is no mention <strong>of</strong> fishes in the<br />

surviving Delacour notebooks (A. Hennache, pers. comm.),<br />

most <strong>of</strong> which disappeared in 1939. Hennache & Dickinson<br />

(2000) provided details <strong>of</strong> Delacour's expeditions, and from<br />

them it appears that Delacour and Lowe collected only once<br />

in Xieng Khouang, Kontum and Dakto before 1936. They<br />

collected at these localities during the second expedition in<br />

1925–1926 (see also Delacour et al., 1927). Additional fish<br />

material collected by Delacour and Lowe is in MNHN but<br />

were collected during the third (1926–1927) and fourth<br />

(1927–1929) expeditions, except for the types <strong>of</strong> Oreoglanis<br />

delacouri (MNHN 1936.31; see Ng & Kottelat, 1999:<br />

379), which must have come from the second expedition.<br />

No species <strong>of</strong> Homaloptera is known from the coastal<br />

streams on the eastern slope <strong>of</strong> the Annam Range and in the<br />

Red River drainage and this seems to eliminate Tonkin from<br />

the list <strong>of</strong> potential localities. Hennache & Dickinson (2000)<br />

mentioned two localities at which Delacour collected while<br />

in Xieng Khouang: Muongsoui [Muang Sui, nowaday Phu<br />

THE RAFFLES BULLETIN OF ZOOLOGY 2012<br />

51<br />

Kut, about 19°40'N 103°E in Nam Ngum drainage, a Mekong<br />

tributary] and Phu Ké, which is not on my maps, apparently<br />

Phu Keng, a hill about 18 km west-northwest <strong>of</strong> Phonsavan<br />

and also in Nam Ngum drainage. I conducted surveys in<br />

Xieng Khouang and the upper Nam Ngum in 1999 and 2012<br />

and have not seen H. smithi or any H. indochinensis-like<br />

fish. I also have not observed this species in northern Laos<br />

in surveys conducted in 1997, 1999 and 2007.<br />

Hennache & Dickinson (2000) did not provide details<br />

on the Kontum and Dakto localities. Both places are in the<br />

Sesan drainage, a tributary <strong>of</strong> the Mekong. The Sesan joins<br />

the Xe Kong shortly before entering the Mekong in Cambodia.<br />

Homaloptera smithi is a common species in the Xe Kong<br />

(pers. obs.). This makes Kontum or Dakto a likely type locality.<br />

7.7.5 Homalopteroides tweediei (Herre, 1940)<br />

Homaloptera tweediei Herre, 1940: 7, pl. 1 (type locality:<br />

Malaysia: Johore: Mawai District, shallow rapid creek,<br />

about 40 miles north <strong>of</strong> Singapore; holotype: CAS-SU<br />

33102, Böhlke, 1953: 40; noun in genitive, indeclinable)<br />

7.7.6 Homalopteroides wassinkii (Bleeker, 1853)<br />

Homaloptera fasciata van Hasselt, 1823: 133, 1824: 377<br />

(nomen nudum; Kottelat, 1987b: 373)<br />

Balitora ocellata Valenciennes, in Cuvier & Valenciennes,<br />

1846: 96 (type locality: Indonesia: Java: Buitenzorg<br />

[Bogor]; syntypes: MNHN 3122 [1] and specimen illustrated<br />

by Kuhl and van Hasselt [reproduced in Roberts,<br />

1993: fig. 27]; secondary junior homonym <strong>of</strong> Homaloptera<br />

ocellata van der Hoeven, 1830: 211 when placed<br />

in Homaloptera by Bleeker, 1860c: 95; adjective, -us,<br />

-a, -um)<br />

Homaloptera Wassinkii Bleeker, 1853a: 163 (type locality:<br />

Indonesia: Java: Tjampea [Ciampea] and Buitenzorg<br />

[Bogor]; lectotype: RMNH 4987, designated by Alfred,<br />

1961: 36; noun in genitive, indeclinable)<br />

Homaloptera valenciennesi Bleeker, 1860c: 95 (replacement<br />

name for Homaloptera ocellata Valenciennes, in Cuvier<br />

& Valenciennes, 1846: 96; noun in genitive, indeclinable)<br />

Homaloptera fasciata Bleeker, 1860c: 96 (unnecessary replacement<br />

name for H. wassinkii Bleeker, 1853a: 163;<br />

adjective, -us, -a, -um)<br />

Nomenclatural notes. Roberts (1993: 25) considered specimen<br />

MNHN 3122 as the holotype <strong>of</strong> Balitora ocellata. Valenciennes<br />

based his description on a single specimen and<br />

on the drawing sent by Kuhl and van Hasselt, as is obvious<br />

from the mention <strong>of</strong> the colour in life (p. 97) and explicit in<br />

the comment at the end <strong>of</strong> the description <strong>of</strong> B. pavonina (p.<br />

99). Unless it can be demonstrated that the specimen examined<br />

by Valenciennes is the same as figured by Kuhl and van<br />

Hasselt, there is no holotype but a series <strong>of</strong> syntypes.<br />

7.8 Homalopterula Fowler, 1940<br />

Homalopterula Fowler, 1940: 379 (type species: Homalopterula<br />

ripleyi Fowler, 1940: 379, by original designation).<br />

Gender feminine.

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